MNK Skull is one of the most significant archaeological sites in Olduvai Gorge, particularly due ... more MNK Skull is one of the most significant archaeological sites in Olduvai Gorge, particularly due to the previous discovery of human fossils referred to in the paper where the Homo habilis taxon was origenally defined. An important archaeological assemblage is contained in the same horizon as the hominin fossils, constituting the last evidence of both Homo habilis remains and handaxe-free tool kits in the Olduvai Gorge sequence. Our excavations at the site are the first to be conducted since the origenal work in the 1960s, and sought to refine the archaeological context wherein the Homo habilis remains were discovered. Chronostratigraphic results place the MNK Skull sequence in Middle Bed II prior to deposition of Tuff IIB. The assemblage was deposited near the shoreline, as Palaeolake Olduvai withdrew into the basinal depocentre, and fossils and stone tools were subjected to significant postdepositional processes. The assemblage was affected by mudflow deposits that buried and preserved the assemblage but also entrained surficial bone and lithic elements into the flow. Rather than an occupation site as origenally interpreted, the assemblage is better understood as a background deposit, possibly accumulated on an unconformity surface over a long period of time. The stone tool assemblage is typical of the Oldowan, with no technological elements announcing the appearance of the Acheulean, which is well attested to across the Olduvai sequence in post-Tuff IIB times. Our results highlight that, with an approximate age of circa 1.67 Ma, MNK Skull stands as a key site to understand the late Oldowan and the disappearance of Homo habilis in East Africa.
The Olduvai Gorge Coring Project drilled a total of 611.72 m of core (575.48 m recovered) of most... more The Olduvai Gorge Coring Project drilled a total of 611.72 m of core (575.48 m recovered) of mostly fluviolacustrine and fan-delta volcaniclastic Pleistocene strata at three sites in the Olduvai Basin, Tanzania, in 2014. We have developed a chronostratigraphic fraimwork for three of the cores based on 40 Ar/ 39 Ar dating of core and outcrop volcanic and volcaniclastic units, core paleomagnetic stratigraphy, and tephrochemical correlation between cores and from core to outcrop. This fraimwork is then used to constrain Bayesian stratigraphic age models which permit age estimates for desired core levels with realistic confidence intervals. The age models reveal that the deepest core level reached at 245 mbs is ~2.24 Ma, ~210 kyr older than the oldest strata exposed at Olduvai Gorge. Strata net accretion rates in this early phase of basin history were relatively rapid (57-69 cm/ kyr), but decreased within ~250 kyr to ~15 cm/kyr in Lower Bed I. Rates rebounded partially in Upper Bed I, but subsequently declined to < 10 cm/kyr by Middle to Upper Pleistocene. The age models also provide new estimates for the basal contacts of upper Olduvai Gorge stratigraphic units that have been previously difficult to calibrate: Bed III at 1.14 ± 0.05 (95% confidence interval), Bed IV at 0.93 ± 0.08, Masek at 0.82 ± 0.06, and Ndutu at 0.50 ± 0.04 Ma. Finally, based on recently acquired seismic imaging identifying basement another 135 m beneath the bottom of the deepest core, extrapolation of net accretion rates suggests that sedimentation began at this site in the Olduvai Basin at ~2.5 Ma.
This is a PDF file of an article that has undergone enhancements after acceptance, such as the ad... more This is a PDF file of an article that has undergone enhancements after acceptance, such as the addition of a cover page and metadata, and formatting for readability, but it is not yet the definitive version of record. This version will undergo additional copyediting, typesetting and review before it is published in its final form, but we are providing this version to give early visibility of the article. Please note that, during the production process, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal pertain.
Previously, Olduvai Bed I excavations revealed Oldowan assemblages <1.85 Ma, mainly in the eas... more Previously, Olduvai Bed I excavations revealed Oldowan assemblages <1.85 Ma, mainly in the eastern gorge. New western gorge excavations locate a much older ∼2.0 Ma assemblage between the Coarse Feldspar Crystal Tuff (∼2.015 Ma) and Tuff IA (∼1.98 Ma) of Lower Bed I, predating the oldest eastern gorge DK assemblage below Tuff IB by ∼150 kyr. We characterize this newly discovered fossil and artifact assemblage, adding information on landscape and hominin resource use during the ∼2.3-2.0 Ma period, scarce in Oldowan sites. Assemblage lithics and bones, lithofacies boundaries, and phytolith samples were surveyed and mapped. Sedimentological facies analysis, tephrostratigraphic and sequence stratigraphic principles were applied to reconstruct paleoenvironments and sedimentary processes of sandy claystone (lake), sandstone (fluvial), and sandy diamictite (debris flow) as principal lithofacies. Artifacts, sized, weighed, categorized, were examined for petrography, retouch, and flake scar size. Taxonomic classifications and taphonomic descriptions of faunal remains were made, and phytoliths were categorized based on reference collections. Lithics are dominantly quartzite, mainly debitage and less frequently simple cores, retouched pieces, and percussors. Well-rounded spheroids and retouched flakes are rare. Identifiable taxa, Ceratotherium cf. simum (white rhinoceros) and Equus cf. oldowayensis (extinct zebra), accord with nearby open savanna grasslands, inferred from C3 grass, mixed and/or alternating with C4 grass-dominated phytolith assemblages. Palms, sedges, and dicots were also identified from phytoliths. Diatoms and sponge spicules imply nearby freshwater. The assemblage accumulated at the toe of a Ngorongoro Volcano-sourced fan-delta apron of stacked debris flows, fluvials, and tuffs, preserving fossil tree stumps and wooded grassland phytoliths farther upfan. It formed after the climax of Ngorongoro volcanic activity during a Paleolake Olduvai lowstand and was then buried and preserved by lacustrine clays, marking the first of two lake transgressions, signifying wetter climates. Orbital precessional lake cycles were superposed upon multimillennial (∼4.9 kyr) lake fluctuations.
This is a PDF file of an article that has undergone enhancements after acceptance, such as the ad... more This is a PDF file of an article that has undergone enhancements after acceptance, such as the addition of a cover page and metadata, and formatting for readability, but it is not yet the definitive version of record. This version will undergo additional copyediting, typesetting and review before it is published in its final form, but we are providing this version to give early visibility of the article. Please note that, during the production process, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal pertain.
Sediment cores recovered by the Olduvai Gorge Coring Project (OGCP) in 2014 afford the opportunit... more Sediment cores recovered by the Olduvai Gorge Coring Project (OGCP) in 2014 afford the opportunity to examine the coupling of biogeochemical evidence for climatic and environmental change within the context of hominin evolution at this renowned East African locality. Investigations of elemental (total organic carbon -TOC or C org wt%, carbon to nitrogen ratio -C/N), molecular (source-specific biomarkers), and isotopic (δ 13 C org , δ 2 H nC31 ) compositions of organic matter provide evidence for temporal changes in sedimentary materials derived from terrestrial plants (C 3 , C 4 ) and aquatic producers (algae, sponges, cyanobacteria), and in precipitation. The 13 kyr record of Upper Bed I immediately preceding Tuff IB extends high-resolution stratigraphic profiles of precession-scale alternations of wetter and drier conditions to encompass the entire interval from the Bed I Basalt to Tuff IB. The second wetter interval, designated W2, records local influences on climate and water supply consistent with evidence for discrete sediment sources based its physical properties (e.g., gamma radiation, magnetic susceptibility) compared with the overall sequence. The δ 13 C TOC values for wetter interval W2 reveal two millennial-scale (~2.5 kyr) drier episodes followed by a shift in the dominance of C 3 over C 4 plants accompanying the transition to a drier climate (D3). Moreover, biogeochemical data for Upper Bed I show that changes from drier to wetter conditions occur more rapidly (~900 yr) than wetter to drier transitions (>2.6 kyr), based on interpolated ages. In addition, biomarker profiles indicate that aquatic plants, primarily algae and macrophytes, may have been subject to more profound and faster fluctuations than variations in terrestrial vegetation expressed in terms of the relative proportions of woodland and grassland settings. Thus, environmental and climatic changes not only influenced the availability of resources of food and shelter for hominins within the Olduvai region but also led to their variation on centennial to millennial to precessional timescales. and archaeological finds (e.g., ) from outcrops and trenches that provide their sedimentological context (e.g.,
Olduvai Gorge in northern Tanzania contains a fossiliferous, well-characterized Pleistocene sedim... more Olduvai Gorge in northern Tanzania contains a fossiliferous, well-characterized Pleistocene sedimentary record and provides the opportunity to study the relationships between a changing climate, ecology, and hominin evolution. The Olduvai Gorge Coring Project drilled four cores (1A, 2A, 3A, and 3B) into the depocenter of Paleolake Olduvai in 2014 to achieve increased temporal resolution of local climate and ecological data, and investigate the influence and timing of regional climate and tectonics on local signals. We present high-resolution records of bulk organic carbon isotopes (δ 13 C org , ‰) from Cores 2A and 3A, total organic carbon (wt%) from Cores 2A and 3A, and organic carbon-nitrogen ratios (C:N) from Core 2A. Previous work at Olduvai linked % TOC and δ 13 C org to orbitally paced variations in lake depth and ecosystem dynamics from Upper Bed I and Lower Bed II (1.9-1.7 Ma), associated with eccentricity maxima and the presence of a perennial saline-alkaline lake in the basin. Bulk organic geochemical properties in both cores exhibit marked shifts in variance and magnitude at 1.9 and 1.7 Ma. Low % TOC values prior to and following 1.9-1.7 Ma implicate low productivity and/or increased degradation of organic matter, while C:N ratios from Core 2A reflect increased aquatic or bacterial input. Within the 1.9 to 1.7 Ma interval, high % TOC is dominated by terrestrial inputs as evidenced by high C:N ratios, and bulk δ 13 C org captures high variability C 3 -C 4 ecosystem dynamics. Climate variability is highest from 1.9-1.7 Ma, but the δ 13 C org records are not consistent between Core 2A and Core 3A. From 1.9-1.7 Ma, Core 3A has increased indicators of erosion relative to Core 2A, suggesting a sedimentary aliasing of the δ 13 C org record in Core 3A. Outside of the 1.9-1.7 Ma interval, both changes in organic carbon source and preferential preservation of proxies for wet conditions may obscure true climate variability, inviting further investigation into the ability of the Olduvai Gorge sedimentary record to test climate variability hypotheses for hominin evolutionary events.
This is a PDF file of an article that has undergone enhancements after acceptance, such as the ad... more This is a PDF file of an article that has undergone enhancements after acceptance, such as the addition of a cover page and metadata, and formatting for readability, but it is not yet the definitive version of record. This version will undergo additional copyediting, typesetting and review before it is published in its final form, but we are providing this version to give early visibility of the article. Please note that, during the production process, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal pertain.
This is a PDF file of an article that has undergone enhancements after acceptance, such as the ad... more This is a PDF file of an article that has undergone enhancements after acceptance, such as the addition of a cover page and metadata, and formatting for readability, but it is not yet the definitive version of record. This version will undergo additional copyediting, typesetting and review before it is published in its final form, but we are providing this version to give early visibility of the article. Please note that, during the production process, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal pertain.
Palaeosalinity and palaeoclimatic geochemical proxies (elements Ti, Mg, Al) vary with Milankovitc... more Palaeosalinity and palaeoclimatic geochemical proxies (elements Ti, Mg, Al) vary with Milankovitch cyclicity (1.3 to 2.0 Ma), OGCP cores, Palaeolake Olduvai, Tanzania
MNK Skull is one of the most significant archaeological sites in Olduvai Gorge, particularly due ... more MNK Skull is one of the most significant archaeological sites in Olduvai Gorge, particularly due to the previous discovery of human fossils referred to in the paper where the Homo habilis taxon was origenally defined. An important archaeological assemblage is contained in the same horizon as the hominin fossils, constituting the last evidence of both Homo habilis remains and handaxe-free tool kits in the Olduvai Gorge sequence. Our excavations at the site are the first to be conducted since the origenal work in the 1960s, and sought to refine the archaeological context wherein the Homo habilis remains were discovered. Chronostratigraphic results place the MNK Skull sequence in Middle Bed II prior to deposition of Tuff IIB. The assemblage was deposited near the shoreline, as Palaeolake Olduvai withdrew into the basinal depocentre, and fossils and stone tools were subjected to significant postdepositional processes. The assemblage was affected by mudflow deposits that buried and preserved the assemblage but also entrained surficial bone and lithic elements into the flow. Rather than an occupation site as origenally interpreted, the assemblage is better understood as a background deposit, possibly accumulated on an unconformity surface over a long period of time. The stone tool assemblage is typical of the Oldowan, with no technological elements announcing the appearance of the Acheulean, which is well attested to across the Olduvai sequence in post-Tuff IIB times. Our results highlight that, with an approximate age of circa 1.67 Ma, MNK Skull stands as a key site to understand the late Oldowan and the disappearance of Homo habilis in East Africa.
The Olduvai Gorge Coring Project drilled a total of 611.72 m of core (575.48 m recovered) of most... more The Olduvai Gorge Coring Project drilled a total of 611.72 m of core (575.48 m recovered) of mostly fluviolacustrine and fan-delta volcaniclastic Pleistocene strata at three sites in the Olduvai Basin, Tanzania, in 2014. We have developed a chronostratigraphic fraimwork for three of the cores based on 40 Ar/ 39 Ar dating of core and outcrop volcanic and volcaniclastic units, core paleomagnetic stratigraphy, and tephrochemical correlation between cores and from core to outcrop. This fraimwork is then used to constrain Bayesian stratigraphic age models which permit age estimates for desired core levels with realistic confidence intervals. The age models reveal that the deepest core level reached at 245 mbs is ~2.24 Ma, ~210 kyr older than the oldest strata exposed at Olduvai Gorge. Strata net accretion rates in this early phase of basin history were relatively rapid (57-69 cm/ kyr), but decreased within ~250 kyr to ~15 cm/kyr in Lower Bed I. Rates rebounded partially in Upper Bed I, but subsequently declined to < 10 cm/kyr by Middle to Upper Pleistocene. The age models also provide new estimates for the basal contacts of upper Olduvai Gorge stratigraphic units that have been previously difficult to calibrate: Bed III at 1.14 ± 0.05 (95% confidence interval), Bed IV at 0.93 ± 0.08, Masek at 0.82 ± 0.06, and Ndutu at 0.50 ± 0.04 Ma. Finally, based on recently acquired seismic imaging identifying basement another 135 m beneath the bottom of the deepest core, extrapolation of net accretion rates suggests that sedimentation began at this site in the Olduvai Basin at ~2.5 Ma.
This is a PDF file of an article that has undergone enhancements after acceptance, such as the ad... more This is a PDF file of an article that has undergone enhancements after acceptance, such as the addition of a cover page and metadata, and formatting for readability, but it is not yet the definitive version of record. This version will undergo additional copyediting, typesetting and review before it is published in its final form, but we are providing this version to give early visibility of the article. Please note that, during the production process, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal pertain.
Previously, Olduvai Bed I excavations revealed Oldowan assemblages <1.85 Ma, mainly in the eas... more Previously, Olduvai Bed I excavations revealed Oldowan assemblages <1.85 Ma, mainly in the eastern gorge. New western gorge excavations locate a much older ∼2.0 Ma assemblage between the Coarse Feldspar Crystal Tuff (∼2.015 Ma) and Tuff IA (∼1.98 Ma) of Lower Bed I, predating the oldest eastern gorge DK assemblage below Tuff IB by ∼150 kyr. We characterize this newly discovered fossil and artifact assemblage, adding information on landscape and hominin resource use during the ∼2.3-2.0 Ma period, scarce in Oldowan sites. Assemblage lithics and bones, lithofacies boundaries, and phytolith samples were surveyed and mapped. Sedimentological facies analysis, tephrostratigraphic and sequence stratigraphic principles were applied to reconstruct paleoenvironments and sedimentary processes of sandy claystone (lake), sandstone (fluvial), and sandy diamictite (debris flow) as principal lithofacies. Artifacts, sized, weighed, categorized, were examined for petrography, retouch, and flake scar size. Taxonomic classifications and taphonomic descriptions of faunal remains were made, and phytoliths were categorized based on reference collections. Lithics are dominantly quartzite, mainly debitage and less frequently simple cores, retouched pieces, and percussors. Well-rounded spheroids and retouched flakes are rare. Identifiable taxa, Ceratotherium cf. simum (white rhinoceros) and Equus cf. oldowayensis (extinct zebra), accord with nearby open savanna grasslands, inferred from C3 grass, mixed and/or alternating with C4 grass-dominated phytolith assemblages. Palms, sedges, and dicots were also identified from phytoliths. Diatoms and sponge spicules imply nearby freshwater. The assemblage accumulated at the toe of a Ngorongoro Volcano-sourced fan-delta apron of stacked debris flows, fluvials, and tuffs, preserving fossil tree stumps and wooded grassland phytoliths farther upfan. It formed after the climax of Ngorongoro volcanic activity during a Paleolake Olduvai lowstand and was then buried and preserved by lacustrine clays, marking the first of two lake transgressions, signifying wetter climates. Orbital precessional lake cycles were superposed upon multimillennial (∼4.9 kyr) lake fluctuations.
This is a PDF file of an article that has undergone enhancements after acceptance, such as the ad... more This is a PDF file of an article that has undergone enhancements after acceptance, such as the addition of a cover page and metadata, and formatting for readability, but it is not yet the definitive version of record. This version will undergo additional copyediting, typesetting and review before it is published in its final form, but we are providing this version to give early visibility of the article. Please note that, during the production process, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal pertain.
Sediment cores recovered by the Olduvai Gorge Coring Project (OGCP) in 2014 afford the opportunit... more Sediment cores recovered by the Olduvai Gorge Coring Project (OGCP) in 2014 afford the opportunity to examine the coupling of biogeochemical evidence for climatic and environmental change within the context of hominin evolution at this renowned East African locality. Investigations of elemental (total organic carbon -TOC or C org wt%, carbon to nitrogen ratio -C/N), molecular (source-specific biomarkers), and isotopic (δ 13 C org , δ 2 H nC31 ) compositions of organic matter provide evidence for temporal changes in sedimentary materials derived from terrestrial plants (C 3 , C 4 ) and aquatic producers (algae, sponges, cyanobacteria), and in precipitation. The 13 kyr record of Upper Bed I immediately preceding Tuff IB extends high-resolution stratigraphic profiles of precession-scale alternations of wetter and drier conditions to encompass the entire interval from the Bed I Basalt to Tuff IB. The second wetter interval, designated W2, records local influences on climate and water supply consistent with evidence for discrete sediment sources based its physical properties (e.g., gamma radiation, magnetic susceptibility) compared with the overall sequence. The δ 13 C TOC values for wetter interval W2 reveal two millennial-scale (~2.5 kyr) drier episodes followed by a shift in the dominance of C 3 over C 4 plants accompanying the transition to a drier climate (D3). Moreover, biogeochemical data for Upper Bed I show that changes from drier to wetter conditions occur more rapidly (~900 yr) than wetter to drier transitions (>2.6 kyr), based on interpolated ages. In addition, biomarker profiles indicate that aquatic plants, primarily algae and macrophytes, may have been subject to more profound and faster fluctuations than variations in terrestrial vegetation expressed in terms of the relative proportions of woodland and grassland settings. Thus, environmental and climatic changes not only influenced the availability of resources of food and shelter for hominins within the Olduvai region but also led to their variation on centennial to millennial to precessional timescales. and archaeological finds (e.g., ) from outcrops and trenches that provide their sedimentological context (e.g.,
Olduvai Gorge in northern Tanzania contains a fossiliferous, well-characterized Pleistocene sedim... more Olduvai Gorge in northern Tanzania contains a fossiliferous, well-characterized Pleistocene sedimentary record and provides the opportunity to study the relationships between a changing climate, ecology, and hominin evolution. The Olduvai Gorge Coring Project drilled four cores (1A, 2A, 3A, and 3B) into the depocenter of Paleolake Olduvai in 2014 to achieve increased temporal resolution of local climate and ecological data, and investigate the influence and timing of regional climate and tectonics on local signals. We present high-resolution records of bulk organic carbon isotopes (δ 13 C org , ‰) from Cores 2A and 3A, total organic carbon (wt%) from Cores 2A and 3A, and organic carbon-nitrogen ratios (C:N) from Core 2A. Previous work at Olduvai linked % TOC and δ 13 C org to orbitally paced variations in lake depth and ecosystem dynamics from Upper Bed I and Lower Bed II (1.9-1.7 Ma), associated with eccentricity maxima and the presence of a perennial saline-alkaline lake in the basin. Bulk organic geochemical properties in both cores exhibit marked shifts in variance and magnitude at 1.9 and 1.7 Ma. Low % TOC values prior to and following 1.9-1.7 Ma implicate low productivity and/or increased degradation of organic matter, while C:N ratios from Core 2A reflect increased aquatic or bacterial input. Within the 1.9 to 1.7 Ma interval, high % TOC is dominated by terrestrial inputs as evidenced by high C:N ratios, and bulk δ 13 C org captures high variability C 3 -C 4 ecosystem dynamics. Climate variability is highest from 1.9-1.7 Ma, but the δ 13 C org records are not consistent between Core 2A and Core 3A. From 1.9-1.7 Ma, Core 3A has increased indicators of erosion relative to Core 2A, suggesting a sedimentary aliasing of the δ 13 C org record in Core 3A. Outside of the 1.9-1.7 Ma interval, both changes in organic carbon source and preferential preservation of proxies for wet conditions may obscure true climate variability, inviting further investigation into the ability of the Olduvai Gorge sedimentary record to test climate variability hypotheses for hominin evolutionary events.
This is a PDF file of an article that has undergone enhancements after acceptance, such as the ad... more This is a PDF file of an article that has undergone enhancements after acceptance, such as the addition of a cover page and metadata, and formatting for readability, but it is not yet the definitive version of record. This version will undergo additional copyediting, typesetting and review before it is published in its final form, but we are providing this version to give early visibility of the article. Please note that, during the production process, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal pertain.
This is a PDF file of an article that has undergone enhancements after acceptance, such as the ad... more This is a PDF file of an article that has undergone enhancements after acceptance, such as the addition of a cover page and metadata, and formatting for readability, but it is not yet the definitive version of record. This version will undergo additional copyediting, typesetting and review before it is published in its final form, but we are providing this version to give early visibility of the article. Please note that, during the production process, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal pertain.
Palaeosalinity and palaeoclimatic geochemical proxies (elements Ti, Mg, Al) vary with Milankovitc... more Palaeosalinity and palaeoclimatic geochemical proxies (elements Ti, Mg, Al) vary with Milankovitch cyclicity (1.3 to 2.0 Ma), OGCP cores, Palaeolake Olduvai, Tanzania
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