Climax

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Climax

• Culmination of a directional change is a steady-state, a climax


community, in which no further directional change takes place under the
prevailing environmental conditions.
• It is the terminus of habitat and vegetation development.
• The climax community is a steady state with respect to productivity,
structure and population.
• The climax community has a maximum diversity, relative stability and
homogeneity of the spp. populations within and between stands.
• Each stand is self-maintaining and long-persisting.
• The composition and population of the climax are determined by all
factors of a mature ecosystem like
o The properties of each species
o Biotic inter-relations
o Floristic & Faunistic availability
o Chances of dispersal
o Inter-specific and intra-specific interaction
o Type of soil
o Climatic situation
o And special factors like fire, wind, snow and salinity.
• There is a mosaic of climax types, corresponding to the mosaic of
habitats or environmental gradients (Hanson & Churchill, 1961).
• However, changes are expected within the climax and fluctuation change.

THEORIES OF CLIMAX

1. Monoclimax Concept (Clements, 1916)


• Only one true climax community is possible, which is mainly controlled
by the climatic factors.
• It is thus a climatic climax
• It is not affected by soil or topography
• Subordinate communities are: proclimax, preclimax, postclimax,
sereclimax, subclimax and disclimax
Proclimax: Another stable community developing on the margins of climax
community. On disturbance, it can easily be changed in appearance.
Preclimax: When climax is localized in regions or margins in which plants are of
lower order than climax. It is often a drier patch or region for example
on a higher elevation on a mountain slope.

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Postclimax: It is often developed in regions having greater moisture than in normal
climax community. A strip of vegetation of higher life-forms found
within a climax e.g. a forest along a stream in grassland. Another
example is a community at a lower altitude than a particular climax
community, e.g. on a mountain. An example is of Potohar Plateau in
the plains of Punjab.
Sereclimax: The community, which becomes stabilized at any of the seral stages of
succession due to microclimate.
Subclimax: The stage in succession just preceding the climatic climax
Disclimax: The community, which becomes stabilized due of recurrent
disturbance by man or other biotic factors.
• According to the monoclimax concept, all these communities, over a long
period of time are believed to undergo modification by the action of
climate into a true climax vegetation

2. Polyclimax Concept: (Tansley, 1935)


• A climax reflects not only the climatic factors but also other factors of
the environmental complex i.e. edaphic, biotic etc.
• The climax can be of several other kinds, which are different from the
climatic climax of the area.
• Climate is not the only factor that determines the climax in a large area,
other factors like edaphic, topographic and biotic factors are equally
important.
• Thus, within the same climatic zone, edaphic, topographic and biotic
climaxes may occur in localized regions.
• In this way in addition to the climatic climax, different climax stages may
develop e.g. Edaphic climax, Topographic climax, Biotic climax
(Disclimax), Fire climax etc.

3. Climax Pattern Theory: (Whittaker, 1953)


• The stable plant communities are developed by the effect of all the
environmental factors at a particular place.
• Vegetation may not be divided into small sub-units.
• The most important point of this theory is that only one large community
develops according to soil, slope and other habitat factors.
• Various climax types are orderly organized due to different
environmental conditions.

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