lg401.v07 Phytoplankton Analysis - Rfa

Standard Operating Procedure
for Phytoplankton Analysis
LG401
Version 07, March 2021

TABLE OF CONTENTS
Section
Number Subject Page
1.0 SCOPE AND APPLICATION ............................................................................................................ 1

2.0 SUMMARY OF METHOD................................................................................................................. 1
3.0 SAMPLE COLLECTION ................................................................................................................... 1
4.0 APPARATUS ...................................................................................................................................... 1
5.0 REAGENTS ........................................................................................................................................ 2
6.0 ANALYTICAL PROCEDURE ........................................................................................................... 2
7.0 ARCHIVING ....................................................................................................................................... 8
8.0 CALCULATIONS ............................................................................................................................... 9
9.0 QUALITY CONTROL AUDITS AND METHOD PRECISION ....................................................... 9
Appendix 1: Geometric Forms and the Measurements Required ........................................................................ 11
Figure 1: Image of Ceratium ................................................................................................................................ 12
Appendix 2: Taxonomy and Shape Codes for Great Lakes Phytoplankton Taxa................................................ 13
Appendix 3: Formulas for Calculating Biomass for Various Phytoplankton Shapes .......................................... 35
Disclaimer: Mention of trade names or commercial products does not constitute endorsement or
recommendation for use.
LG401, Version 07, March 2021 ii

Standard Operating Procedure for Phytoplankton Analysis

1.0 SCOPE AND APPLICATION
1.1 This method, as first described by Utermöhl (1958), is utilized to identify and enumerate the
phytoplankton community from many different types of aquatic habitats.
2.0 SUMMARY OF METHOD
2.1 The method, called the Modified Utermöhl method, involves the microscopic examination of a
preserved water sample. Initially a preliminary scan is made to determine the volume of sample
needed for other portions of the procedure. A settled sample of appropriate volume is then
examined for non-diatom algae and Urosolenia species (hereafter referred to as ‘soft algae’). A
second examination is performed on a cleaned diatom preparation for identification and
enumeration.
3.0 SAMPLE COLLECTION
3.1 See U.S. EPA GLNPO Standard Operating Procedure for Phytoplankton Sample Collection
and Preservation Field Procedures (LG400), accessible by logging into glnpo.net and then
going to R/V Lake Guardian SOP Repository.
3.2 A composite (integrated) sample is prepared from the upper region of the water column. For an
unstratified water column, the integrated sample is prepared by taking equal volumes of water from
SRF (1-2 m), 5 m, 10 m and 20 meters unless the depth is less than 20 meters. If the total depth is
between 15 and 22 meters, the 20 meter sample is replaced by the bottom sample (B-1 or B-2). If
the total depth is less than 15 meters, equal volumes are taken from surface, mid-depth, and bottom
sample (B-1 or B-2).
For a stratified water column, equal volumes are taken from the surface, 5 m, 10 m, and lower
epilimnion (LEP). If the epilimnion is very shallow, equal volumes are taken from a maximum of
four sampling depths and a minimum of two sampling depths. The underlying strategy is to collect
a representative sample from the epilimnion.
For more detailed instructions on depth selection for the integrated sample, see Field Sampling Using
the Rosette Sampler (LG200), accessible by logging into glnpo.net and then going to R/V Lake
Guardian SOP Repository.
4.0 APPARATUS
4.1 Inverted microscope with an objective system for magnification up to 600x (Leitz Diavert or
another equal quality inverted microscope)
4.2 Compound microscope with an objective system of magnification of 1000x or greater
4.3 Sedimentation chambers: 5-, 10-, 25-, 50- and 100-cc
LG401, Version 07, March 2021 Page 1 of 35

4.4 Beakers: 300- and 600-mL
4.5 Large hotplate capable of boiling water
4.6 Centrifuge
4.7 Centrifuge tubes, graduated 50-mL
4.8 Coverslips, round, #1 thickness, 22-mm diameter
4.9 Pre-cleaned microscope slides, 25 x 75 mm
4.10 Syringe, 20-mL with cannula, 14-gauge 4-inch (optional for strewing, or use macropipetter)
4.11 Long-neck disposable pipettes or macropipetter
4.12 Rubber bulbs for pipettes (if disposable pipettes used)
4.13 10-mL autopipette
5.0 REAGENTS
5.1 HNO3 = Nitric Acid (concentrated)
5.2 H2O2 = Hydrogen peroxide (30% solution)
5.3 K2Cr2O7 = Potassium dichromate
5.4 HyraxTM mounting media, or equivalent (Naphrax, Pleurax, etc.)
5.5 Toluene/xylene
5.6 Commercial formaldehyde solution 37 - 40% (= formalin)
5.7 Immersion oil for upright microscopy
6.0 ANALYTICAL PROCEDURE
6.1 Phytoplankton samples received at the lab shall be logged-in and spiked with 10 ml of Formalin.
6.2 10-mL Preliminary Investigation
6.2.1 An initial screening of each sample must be done in order to determine the final settled
volume needed for analyses unless historical data is available to show what volumes have
traditionally been used for samples from the same site. This is done by sedimenting 10 mL
of each sample and counting the total number of photosynthetic organisms, and the number

of diatom cells, within a select area of the slide (10 mm2 minimum or 3 transects of the
chamber). No identifications are done at this time, but any irregularities such as excessive
sediment in the sample are noted.
The volume needed for settling and subsequent soft algae analysis is determined from the
number of all organisms counted. The sample volume to be digested for diatom analysis is
determined from the preliminary diatom cell counts. However, the minimum volume for
digestion is recommended to be 500 mL. For example:
10-mL preliminary counts
1) 101 organisms total

2) 103 diatom cells (note: 1 cell has 2 frustules/valves)
Count needed (minimum)
1) 250 organisms total

2) 500 diatom frustules (250 cells)
Final volumes
1) 25-mL sample for sedimentation

2) 500-mL sample for digestion
The final volume may be slightly over-estimated to ensure that the minimum counts required
are met. The preliminary count also helps to ensure that there is enough sample for both final
investigations.
The definition of an organism is as follows:
A colony, a filament, or a single cell. The units of a colony or a filament are not
counted as organisms at this time but the whole aggregate is counted as one
organism.
6.2.2 All information from the 10-mL preliminary count is recorded in a pre-printed data form,
generally on the first page of the countsheet used for soft algae analysis. This includes
unusual observations such as poor sample preservation, high bacterial/fungal populations,
occurrence of special/rare phytoplankton taxa, and the degree of matrix interference, etc.
6.3 Sample Sedimentation
6.3.1 The phytoplankton sample is homogenized by gently inverting the sample bottle for 60
seconds. The predetermined sample volume is loaded into a settling Utermöhl chamber of
appropriate volume. Samples should be added to the chamber with a syringe and/or
macropipettor with a clean tip. The sample bottle should be inverted at least once between
each addition. This is done because larger organisms settle quickly and may remain in the
bottle if the sample is simply poured. The chamber is topped with a round top plate.

6.3.2 Algae are allowed to settle onto the base of the settling chamber. Since oil immersion may
be used in the course of identification, the coverglass at the bottom of the chamber should
not be thicker than 0.2 - 0.3 mm inches (or No. 1 coverglass). The time recommended for
complete sedimentation varies with the height of the chamber (8 cm/day to 4 cm/day
depending on accuracy required in enumeration (Furet & Benson-Evans, 1982)).
6.3.3 Approximate settling times necessary are as follows:
100 mL ----------------- 100 hours

50 mL ------------------ 50 hours
25 mL ------------------ 25 hours
10 mL ------------------ 10 hours
5 mL 5 hours
2 mL 2 hours
6.4 Total Sample Analysis
6.4.1 A complete phytoplankton analysis consists of two parts. The first part is a count of all
organisms in the settled sample at 400-500x. The second part is a count made on a
prepared diatom slide at least 1000x.
6.5 Sedimented Sample Analysis
6.5.1 The soft algae portion of the settled phytoplankton samples are examined and analyzed
using an inverted microscope (Leitz Diavert, Olympus BX51 or equivalent microscope).
6.5.2 All “live” forms (chloroplast containing organisms) are counted and identified at
400x. Higher magnification may be used for identification when necessary.
6.5.3 Procedure
6.5.3.1 The entire chamber of settled material is scanned and the dominant (4 or 5 most
common organisms) as well as subdominant species determined. This is to give the
biologist an idea of the sample composition as well as to ensure that the sample is
evenly settled.
6.5.3.2 A minimum of 250 “live” organisms is counted along transects. The area
counted is recorded since it is needed for cells per mL calculation.
6.5.3.3 For the purposes of determining the numbers of organisms to count, colonies and
filaments are considered one organism. However, for the purposes of calculating
biovolume, individual cells within colonies and filaments are counted and
measured. Where numbers of cells cannot be counted, e.g., in extremely large
cyanobacterial colonies, estimates can be made.

6.5.3.4 Large organisms, such as Ceratium hirundinella, should be enumerated from the
whole chamber. Calculations of cell numbers of such organisms should be done
using the area of the entire chamber bottom.
6.5.3.5 The number of “live” cells are enumerated at the lowest taxonomic unit possible
(i.e., genus, species, variety, etc.). All “empty” lorica from Chrysophyta are also
enumerated and identified to species level where possible, although these are not
included in the ‘regular’ cell counts.
6.5.3.6 As many as 20 specimens of each species are measured for cell volume calculations.
When fewer than 20 specimens are present, those present are measured as they
occur. The measurements required are those which are necessary for the volume
calculation of a solid which best approximates the shape of any particular organism.
For most organisms the measurements are taken from out-side wall to outside wall.
In cases where a taxon’s size is invariable (as for some small green algae and
cyanophytes) it is permissible to record the measurement once.
6.5.3.7 Those forms which are loricate (Chlorophyta: Phacotaceae and Chlorococaceae;
Chrysophyta: many forms) must have the active portion, i.e., protoplast measured.
Filamentous and colonial forms require measurements of individual components. If
cell walls are not readily visible in the filament, the length of the entire colony is
measured and the number of cells is determined by dividing by the average cell
length which is measured when possible.
6.5.3.8 During examination of the settled sample, most diatoms are enumerated and
identified only as live pennates, empty pennates, live centrics, and empty centrics,
with the only exception being species of Urosolenia (=Rhizosolenia). Actual
species identification of diatoms (excluding Urosolenia) and cell volume
measurements are done under oil immersion (1000-1250x) by another method
(Section 6.6). While not included in the regular counts, note should be made of
the presence of other identifiable species, such as Fragilaria capucina, Fragilaria
crotonensis, Tabellaria flocculosa, and Stephanodiscus binderanus, to provide
corroboration of identifications in cases where colonial configuration is a
characteristic feature.
6.6 Diatom Sample Analysis
6.6.1 The cellular contents of diatoms obscure the wall markings on which the taxonomy is based.
Therefore, the organic matter must be removed (oxidized) prior to identification. The
following method describes a cleaning method, slide preparation, enumeration, as well as
identification.
6.6.2 Cleaning of Diatoms
Sample cleaning and slide preparation must be performed in the hood.

6.6.2.1 Homogenize the sample by gently inverting the sample bottle every second for a
minimum of one minute. Pour a specified volume (dependent upon diatom
density and determined by the initial 10-mL count discussed previously) of
homogenized sample is put into a 600-mL (or larger) beaker. Unless diatom
densities are extremely high, a minimum volume of 500 mL should be used.
6.6.2.2 Add 20 mL of concentrated HNO3 to the beaker. Then place the beaker on a
hotplate and heat until volume is reduced to about 50 mL.
6.6.2.3 Pour the remaining sample into a 300-mL beaker. Completely rinse the sides of the
beaker with RO/DI or distilled water at least three times and empty this rinse water
into the beaker. If necessary, repeat this process several times until a volume of 125
mL is reached. Put 25 mL 30% H2O2 into the beaker, and a few grains of crystal
K2Cr2O7. Place sample on hotplate again and heat until volume is reduced to less
than 10-15 mL.
6.6.2.4 After sample is reduced, transfer the sample into a 50-mL graduated centrifuge
tube. Rinse the beaker at least three times with small quantities of distilled water
and add rinse water to thetube to ensure all diatom remains are transferred to the
centrifuge tube. All centrifuge tubes should contain the same volume,
approximately 35 mL, to ensure centrifuge is balanced. Centrifuge at low speed
(2000 rpm) for 15 minutes. Alternatively, samples can be concentrated by settling
in glass test tubes or vials, using the same settling assumptions as shown in Section
6.3.3.
6.6.2.5 Draw off all but 5 mL of supernatant, ensuring that the pellet is not disturbed. Add
distilledwater to 35 mL, gently shake the sample using a vortex mixer, and
centrifuge again for 15 minutes at 2000 rpm. Repeat this step 7 times.
6.6.2.6 Upon final centrifugation draw off all but ~5 mL of supernatant. Bring volume to
approximately 5 mL with RO/DI or distilled water as necessary. This is the
“cleaned” sample to be used to prepare diatom slide for analysis.
6.6.3 Diatom Slide Preparation
6.6.3.1 Two slides are made from each sample; “A” and “B”. The “B” slide is considered a
duplicate.
6.6.3.2 Place a clean coverslip on a slide warmer (not turned on). The slide warming tray
reduces static charge that may come from other surfaces, but the samples are never
warmed; drying coverslips by warming tends to cause clumping of the diatom
remains, so the coverslips are allowed to evaporate at room temperature.
6.6.3.3 Gently homogenize the pellet and pipette about 0.5 mL of the concentrated sample
on the center of a coverslip and let dry. A larger or smaller aliquot may be used
depending on the diatom densities. When dry, observe each coverslip under a
compound microscope to be sure that there is an adequate density of frustules to

allow counting. If there is not a sufficient density for counting, increase frustule
density by adding more sample.
6.6.3.4 When coverslip drying is complete add a drop of mounting medium (Hyrax,
Naphrax, Pleurax or comparable high-refractive-index medium) to the center of a
clean pre-labeled slide (75 x 25 mm). If the mounting medium is too viscous, add a
few drops (1 to 2 mL) of toluene and/or xylene to dilute the medium.
6.6.3.5 Mount the coverslip, diatom side down, on the medium on the slide and place on
hotplate.
6.6.3.6 Allow solvent to evaporate until bubbles are no longer formed under the coverslip.
Remove from hotplate.
6.6.3.7 Press coverslip gently with pencil eraser or tweezers to extrude excess
mounting medium immediately after removing from heat, as the medium sets
up very quickly.
6.6.3.8 Allow slide to cool and remove excess mounting medium before examining. It will
scrape away easily with a razor blade if all of the solvent is removed: if it is sticky,
return to the hotplate to remove any remaining solvent.
6.6.3.9 Clean, label, and store the slide properly. The label should include the sample
number, year, and station.
6.6.4 Diatom Enumeration and Identification
6.6.4.1 Diatoms should be identified and enumerated at 1000-1250x. Identification should

be down to the finest taxonomic rank possible.
6.6.4.2 Count at least 500 frustules (2 frustules = 1 diatom cell) per sample.
6.6.4.3 At least 10 specimens of each species are measured (wall to wall) for cell volume
calculations. When fewer than 10 specimens are present, those present are measured
as they occur. Measurements should be recorded as cell measurements. For example,
when measuring the depth (i.e., length) of Aulacoseira, be sure to either measure
two frustules together, or double the measurement of a single valve. In some
species, e.g. most Stephanodiscus and Cyclotella, depth measurements are
problematic in that cells are seldom oriented in girdle view, and even when they are,
total cell depth is not easily estimable from the depth of a single frustule due to
overlap of the girdles. Care must therefore be taken in making these measurements.
6.6.5 General Analysis Guidelines
6.6.5.1 Unknown species should be referred to as Genus spp. Taxa for which in-house
descriptions exist, but descriptions have not been published, should be given the
appropriate number (e.g., Stephanodiscus #10). It is crucial that these designations

correspond only to previously established taxa. If an analyst encounters a species that

is not on the species list and feels there is sufficient evidence that it should be added
to the list, it must be confirmed by an outside expert and approved by the GLBMP
Technical Lead before it is added to the species list. In the case of ‘soft’ algae, the
organism should be photographed or a drawing should be made, making clear the
distinguishing characteristics of the species, a written description provided, and
reference made to the taxonomic work which contains the key/description that was
used to identify the organism.
For diatoms, in addition to the above, the frustule should be marked on the slide,
using, e.g., a diamond tipped etching tool. These materials should be sent to an
appropriate expert for confirmation, and the results of confirmation, in addition to
the original supporting materials, should be sent to the WAM, along with a brief
memo describing the species and requesting inclusion in the species list. The
WAM will then send a memo back, either approving of disapproving inclusion of
the taxon on the species list.
6.6.5.2 A note on phytoplankton taxonomy. Appendix 2 contains the currently accepted list
of phytoplankton taxa enumerated from Great Lakes samples. To ensure long-term
consistency, this list contains taxonomic nomenclature from earlier versions of this
SOP (columns 3 and 4). Many of these genus and species names are deprecated, so a
corresponding list of contemporary names and authorities is provided. This updated
taxonomy for the flora is considered correct as of October 2019. In some cases we
have noted taxonomy that is actively undergoing transition, such as the ambiguity in
the Cyclotella sensu lato genera Lindavia and Pantocsekiella. Further, some taxa
have been merged (e.g. Gyrosigma nodiferum has been deemed a synonym of
Gyrosigma sciotoense) but we have retained the obsolete names in the list to ensure
older phytoplankton data are correctly assigned when contemporary taxonomy is
used.
7.0 ARCHIVING
7.1 Soft algae
7.1.1 Soft algae samples are to be archived one data set (usually one year) at a time.
7.1.2 Gently homogenize the remainder of the phytoplankton sample by inverting the bottle for
about 1 minute. Carefully empty the sample into a 500-mL graduated cylinder and cover the
cylinder with a plastic Petri plate. Record the volume of sample settled on a pre-printed
phytoplankton archive form. A larger and/or smaller graduated cylinder may be used
depending on the volume remaining in phytoplankton sample bottle.
7.1.3 Rinse the sample bottle three times with a small amount of RO/DI or distilled water (about
5 mL). Empty the rinse water into the graduated cylinder.
7.1.4 Settle the sample for a minimum of 7 days, but not more than 14 days. Do not disturb the
cylinder.

7.1.5 At the end of the settling period, carefully siphon off the top of the water column without
disturbingthe settled materials. Generally, about 18 - 22 mL of the sample should be
remaining in the cylinder.
7.1.6 Decant the remaining sample from the graduated cylinder into a pre-labeled 25-mL glass
liquid scintillation vial. Rinse the cylinder two times with about 2 mL of RO/DI or distilled
water and emptythe rinse water into the vial. This is the archived sample.
7.1.7 Add about 0.5 mL of formalin solution into the vial before putting the cap on the vial.
7.1.8 To minimize evaporation, parafilm, or some other suitable substance, should be wrapped
around the cap.
7.1.9 Store the archived sample in a pre-labeled tray/box.
7.2 Diatoms
7.2.1 After the diatom slides are made, transfer the remainder of “cleaned” sample to a pre-labeled
glass vial.
7.2.2 Store the diatom archived sample in a box for future reference.
8.0 CALCULATIONS
8.1 Report the results of the sample sedimentation procedure as cells per mL, which is calculated as
follows:
Cells/mL = (C * TA) / (L * W * V * S)
where:
C = cell tally
L = length of strip (mm)
W = width of strip (mm)
V = volume of chamber (mL)
S = number of strips counted
TA = total area of chamber bottom (mm2)
In the case of species (e.g., Ceratium hirundinella) for which the entire chamber bottom is
examined, the formula reduces to:
Cells/mL = (C/V)
8.2 Biovolume is calculated using formulas representing the closest approximation of geometric
shape. A list of geometric forms and the measurements required, are listed in the appendices.
8.3 The data from the diatom slides is reported as percent composition of the 1250x count. This
percent is applied back to the live diatom counts at 500x to determine a cells/mL count for each
species.

8.3.1 Calculate the total live centric (excluding Urosolenia) and live pennate diatom cells/mL
as per formula in Section 8.1.
8.3.2 For each diatom species encountered during the slide counts, calculate what percentage
of the total number of centrics or pennates it represents by dividing the number observed
by the total centric or pennate diatom values enumerated.
8.3.3 Multiply this number by the appropriate number calculated in 8.3.1. This is the cells/mL
for that species.
9.0 QUALITY CONTROL AUDITS AND METHOD PRECISION
9.1 Ten percent of all samples collected are analyzed by a second analyst.
9.1.1 Duplicate counts and measurements by two analysts are performed for both Utermöhl
samples and diatom slide counts from a given sample selected for precision analyses.
9.1.2 Utermöhl samples are counted by the second analyst while still in the counting
chamber so that only interanalyst variation is quantified, and not variation associated
with sub-sampling.
9.1.3 Results from the second analyst are reported under the same sample number as the original
sample, with the exception that the seventh character is replaced by a “Q”. For instance,
12GC19I72 becomes 12GC19Q72.
9.1.4 The Bray-Curtis Index is used as a quantitative method of species-level comparison for
both enumerations and calculated biovolumes produced by the two analysts. The Bray-
Curtis measure is calculated as follows:
∑𝑛𝑛𝑛𝑛𝑖𝑖𝑖𝑖=1(𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚𝑚�𝐴𝐴𝐴𝐴𝑖𝑖𝑖𝑖𝑗𝑗𝑗𝑗 , 𝐴𝐴𝐴𝐴𝑖𝑖𝑖𝑖𝑗𝑗𝑗𝑗 �)
𝑃𝑃𝑃𝑃𝑃𝑃𝑃𝑃 = 200
𝑗𝑗𝑗𝑗𝑗𝑗𝑗𝑗
∑𝑛𝑛𝑛𝑛
𝑖𝑖𝑖𝑖=1 𝐴𝐴𝐴𝐴𝑖𝑖𝑖𝑖𝑖𝑖𝑖𝑖 + 𝐴𝐴𝐴𝐴𝑖𝑖𝑖𝑖𝑖𝑖𝑖𝑖
where:
PSjk = percentage similarity between sample analyses j and k,
Aij = abundance of taxon i in sample analysis j,
Aik = abundance of taxon i in sample analysis k, and
n = total number of taxa in sample analyses j and k.
9.1.5 An interim minimum acceptance value of 60% similarity is currently being used until
enough data accumulate to determine a more appropriate value. The two taxonomists
will discuss the results from all samples which fail to meet this criterion. Outcomes of
such discussions are included as part of dataset reporting to the USEPA. If a major
difference is found in how the two analysts have been identifying or measuring
organisms, the last batch of samples that have been counted by the analyst under review
will be recounted or measured.

Appendix 1: Geometric Forms and the Measurements

Required
Dimensions Required
Shape
Code Length Width Depth Diameter
Ceratium CER L W DP D
Cone CON L W
Crucigenia CRU L
Cylinder CYL L W DP D
Dumbell box DBB L W DP
Dumbell DBL L W DP
Diamond Box DMB L W DP
Fusiform FUS L W
Ovoid box OVB L W DP
Ovoid OVO L W
Rectangular box RTB L W DP
Staurastrum STR L W
Tabellaria flocculosa v. geniculata TFG L W DP D
Teardrop TRP L W

Figure 1: Image of Ceratium

Appendix 2: Taxonomy and Shape Codes for Great Lakes Phytoplankton Taxa
2019 UPDATED
2019 UPDATED AUTHORITY
DIVISION SPECCODE GENUS-SPECIES NAME AUTHORITY TAXONOMIC NAME SHAPE
(IF APPLICABLE)
(IF APPLICABLE)
BAC ACYNORM Actinocyclus normanii (Gregory ex Grev.) Hust. CYL_DISK
Actinocyclus normanii f.
BAC ACYNORMS subsalsa (Juhl.-Dannf.) Hust. CYL_DISK
BAC ACYSP Actinocyclus sp. CYL_DISK
BAC ATTZACH Attheya zachariasi Brun. Acanthoceras zachariasii (Brun) Simonsen CYL_TUBE
Aulacoseira agassizii var.
BAC AULAGASM malayensis (Hust.) Simonsen CYL_TUBE
BAC AULAMBI Aulacoseira ambigua (Grunow) Simonsen CYL_TUBE
BAC AULDIST Aulacoseira distans (Ehrenb.) Simonsen CYL_TUBE
Aulacoseira distans var.
BAC AULDISTA alpigena (Grun.) Simonsen Aulacoseira alpigena (Grunow) Krammer CYL_TUBE
Aulacoseira distans var.
BAC AULDISTL (O.Müll.) Simonsen CYL_TUBE
limnetica
BAC AULGRAN Aulacoseira granulata (Ehrenb.) Simonsen CYL_TUBE
Aulacoseira granulata var.
BAC AULGRANA angustissima (O.Müll.) Simonsen CYL_TUBE
Aulacoseira granulata var.
BAC AULGRANV valida (Hust.) Simonsen CYL_TUBE
BAC AULISLA Aulacoseira islandica (O.Müll.) Simonsen CYL_TUBE
BAC AULITAL Aulacoseira italica (Ehrenb.) Simonsen CYL_TUBE
Aulacoseira italica var.
BAC AULITALT (Grunow) Simonsen CYL_TUBE
tenuissima
BAC AULSP Aulacoseira sp. CYL_TUBE
BAC AULSUBA Aulacoseira subarctica (O.Müll.) E.Y.Haw CYL_TUBE
BAC CTOSP Chaetoceros sp. CYL_TUBE
BAC COSLACU Coscinodiscus lacustris Grun. Thalassiosira lacustris (Grunow) Hasle CYL_DISK
Coscinodiscus lacustris var. Thalassiosira hyperborea
BAC COSLACUS septentrionalis Grun. var. septentrionalis (Grunow) Hasle CYL_DISK
BAC COSSPB Coscinodiscus sp. Thalassiosira sp. CYL_DISK
(H.Kobayasi & H.Kobay.)
Cyclostephanos
BAC CYSCOST Stoermer, Håk. & CYL_DISK
costatilimbus E.C.Ther.
BAC CYSDUBI Cyclostephanos dubius (Fricke) Round CYL_DISK
(M.H.Hohn & Hellermann)
BAC CYSINVI Cyclostephanos invisitatus E.C.Ther., Stoermer & CYL_DISK
Håk.
BAC CYSSP Cyclostephanos sp. CYL_DISK
Stoermer, Håk. &
BAC CYSTHOL Cyclostephanos tholiformis CYL_DISK
E.C.Ther.
Lindavia antiqua (likely (W.Sm.) Nakov, Guillory, M.L.Julius,
BAC CYCANTI Cyclotella antiqua W. Sm. Pantocsekiella) E.C.Ther. & A.J.Alverson CYL_DISK
BAC CYCATOM Cyclotella atomus Hust. CYL_DISK
Cyclotella atomus “fine
BAC CYCATOMF form” In-house form CYL_DISK
BAC CYCCATE Cyclotella catenata (Brun) H.Bachm. CYL_DISK
Lindavia comensis (likely (Grunow) Nakov, Guillory, M.L.Julius,
BAC CYCCOMES Cyclotella comensis Grun. CYL_DISK
Pantocsekiella) E.C.Ther. & A.J.Alverson
Cyclotella comensis rough Lindavia delicatula (likely(Hust.) Nakov, Guillory, M.L.Julius,
BAC CYCCOMRC center w/ process In-house taxon Pantocsekiella) E.C.Ther. & A.J.Alverson CYL_DISK
Alexson, Wellard Kelly, Estepp &
BAC CYCCOME1 Cyclotella comensis var. 1 Pantocsekiella laurentiana CYL_DISK
Reavie
Lindavia comta (likely (Ehrenb.) Nakov, Guillory, M.L.Julius,
BAC CYCCOMT Cyclotella comta (Ehr.) Kutz. Pantocsekiella) E.C.Ther. & A.J.Alverson CYL_DISK
Cyclotella comta var. Lindavia rossii (likely (Håk.) Nakov, Guillory, M.L.Julius,
BAC CYCCOMTO (Ehr.) Grun. CYL_DISK
oligactis Pantocsekiella) E.C.Ther. & A.J.Alverson
Reimann, J.C.Lewin &
BAC CYCCRYP Cyclotella cryptica Guillard CYL_DISK
Lindavia delicatula (likely (Hust.) Nakov, Guillory, M.L.Julius,
BAC CYCDELI Cyclotella delicatula Hust. CYL_DISK
Pantocsekiella) E.C.Ther. & A.J.Alverson
BAC CYCGAMM Cyclotella gamma Sovereign CYL_DISK
Lindavia glomerata (likely
BAC CYCGLOM Cyclotella glomerata Bachm. Pantocsekiella) (H.Bachm) Adesalu & M.L.Julius CYL_DISK

2019 UPDATED
(IF APPLICABLE)
(IF APPLICABLE)
BAC CYCKRAM Cyclotella krammeri Håk. CYL_DISK
BAC CYCMENE Cyclotella meneghiniana Kütz. CYL_DISK
Lindavia michiganiana (Skvortzov) Nakov, Guillory,
BAC CYCMICH Cyclotella michiganiana Skv. CYL_DISK
(likely Pantocsekiella) M.L.Julius, E.C.Ther. & A.J.Alverson
Lindavia ocellata (likely (Pant.) Nakov, Guillory, M.L.Julius,
BAC CYCOCEL Cyclotella ocellata Pant. Pantocsekiella) E.C.Ther. and A.J.Alverson CYL_DISK
BAC CYCOPER Cyclotella operculata (Ag.) Kutz. Cyclotella distinguenda Hust. CYL_DISK
Cyclotella operculata var. Cyclotella distinguenda
BAC CYCOPERU Hust. (Hust.) Håk. & J.R.Carter CYL_DISK
unipunctata var. unipunctata
Discostella
BAC CYCPSEU Cyclotella pseudostelligera Hust. pseudostelligera (Hust.) Houk & Klee CYL_DISK
Lindavia rossii (likely (Håk.) Nakov, Guillory, M.L.Julius,
BAC CYCROSSI Cyclotella rossi Hak. Pantocsekiella) E.C.Ther. & A.J.Alverson CYL_DISK
Lindavia socialis (likely
BAC CYCSOCI Cyclotella socialis Schutt (?) Pantocsekiella) (F.Schütt) De Toni & Forti CYL_DISK
BAC CYCSP Cyclotella sp. CYL_DISK
BAC CYCSTEL Cyclotella stelligera (Cl. & Grun.) V.H. Discostella stelligera (Cleve & Grunow) Houk & Klee CYL_DISK
BAC CYCSTRI Cyclotella striata (Kütz.) Grunow CYL_DISK
Lindavia tripartita (likely (Pant.) Nakov, Guillory, M.L.Julius,
BAC CYCTRIP Cyclotella tripartita Pant. Pantocsekiella) E.C.Ther. & A.J.Alverson CYL_DISK
BAC CYCWOLT Cyclotella wolterecki Hust. Discostella woltereckii (Hust.) Houk & Klee CYL_DISK
BAC MELROSE Melosira roseana Rabh. Orthoseira roeseana (Rabenh.) Pfitzer CYL_TUBE
BAC MELSP Melosira sp. CYL_TUBE
BAC MELUNDU Melosira undulata (Ehrenb.) Kütz. CYL_TUBE
BAC MELVARI Melosira varians C.Agardh CYL_TUBE
BAC RHIERIE Rhizosolenia eriensis H.L. Sm. Urosolenia eriensis (H.L.Sm.) Round & R.M.Crawford CYL_TUBE
BAC RHILONG Rhizosolenia longiseta Zach. Urosolenia longiseta (O.Zacharias) Edlund & Stoermer CYL_TUBE
BAC RHISPB Rhizosolenia sp. CYL_TUBE
BAC SKEPOTA Skeletonema potamos (C.I.Weber) Hasle CYL_DISK
BAC SKESP Skeletonema sp. CYL_DISK
BAC STEPHCOS Stephanocostis sp. Genkal & Kosmina 1985 CYL_DISK
BAC STEALPI Stephanodiscus alpinus Hust. CYL_DISK
Stephanodiscus alpinus type
BAC STEALP1 I Hust. CYL_DISK
Stephanodiscus alpinus type
BAC STEALP23 II/III Hust. CYL_DISK
BAC STEBIND Stephanodiscus binderanus (Kütz.) Willi Krieg. CYL_DISK
Stephanodiscus binderanus Stephanodiscus
BAC STEBINDO var. oestrupii (A. Cl.) A. Cl. binderanus var. öestrupi (A.Cleve) A.Cleve CYL_DISK
BAC STECARC Stephanodiscus carconensis Grunow CYL_DISK
Stephanodiscus carconensis Stephanodiscus
BAC STECARCP var. pusilla Grun. klamathensis Houk, Klee & H.Tanaka CYL_DISK
Stephanodiscus Stoermer, Håk. &
BAC STECONSP conspicueporus E.C.Ther. CYL_DISK
Stephanodiscus hantzschii f.
BAC STEHANTH hantzschii Hak. & Stoerm. Stephanodiscus hantzschii Grunow CYL_DISK
Stephanodiscus hantzschii f.
BAC STEHANTT tenuis (Hust.) Håk. & Stoermer CYL_DISK
BAC STEMINUT Stephanodiscus minutulus (Kütz.) Cleve & J.D.Möller CYL_DISK
BAC STENIAG Stephanodiscus niagarae Ehrenb. CYL_DISK
Stephanodiscus niagarae
BAC STENIAGM var. magnifica Fricke CYL_DISK
BAC STEPARV Stephanodiscus parvus Stoermer & Håk. CYL_DISK
BAC STESPB Stephanodiscus sp. CYL_DISK
BAC STESP10 Stephanodiscus sp. #10 in house taxon CYL_DISK
BAC STEPARV Stephanodiscus sp. #21 in house taxon CYL_DISK
BAC STESUBT Stephanodiscus subtilis (Goor) A.Cleve CYL_DISK
Stephanodiscus
BAC STEPSBTR subtransylvanicus Gasse CYL_DISK
BAC THABALT Thalassiosira baltica (Grunow) Ostenf. CYL_DISK
BAC THASP Thalassiosira sp. CYL_DISK
BAC THAWEIS Thalassiosira weisflogii (Grun.) G. Fryx. & Hasle Thalassiosira weissflogii (Grunow) G.A.Fryxell & Hasle CYL_DISK
BAC UNICENT Unidentified Centrales CYL_DISK

2019 UPDATED
(IF APPLICABLE)
(IF APPLICABLE)
BAP ACHAFFI Achnanthes affinis Grun. Achnanthidium affine (Grunow) Czarn. DMB
BAP ACHAMOE Achnanthes amoena Hust. Karayevia amoena (Hust.) Bukht. OVB
Achnanthidium
BAP ACHBIAS Achnanthes biasolettiana (Kutz.) Grun. (Lange-Bert.) Lange-Bert OVB
rosenstockii
BAP ACHBIOR Achnanthes bioreti Germ. Psammothidium bioretii (H.Germ.) Bukht. & Round OVB
BAP ACHBREV Achnanthes brevipes C.Agardh OVB
Achnanthes brevipes var.
BAP ACHBREVI intermedia (Kütz.) Cleve OVB
(Cleve) Kulikovskiy, Lange-Bert. &
BAP ACHCALC Achnanthes calcar Cl. Gliwiczia calcar Witkowski OVB
BAP ACHCLEV Achnanthes clevei Grun. Karayevia clevei (Grunow) Bukht. DMB
Achnanthes clevei var. Karayevia clevei var.
BAP ACHCLEVR Hust. (Hust.) Bukht. DMB
rostrata rostrata
BAP ACHCONS Achnanthes conspicua A. Mayer Platessa conspicua (A.Mayer) Lange-Bert. OVB
BAP ACHDEFL Achnanthes deflexa Reim. in Patr. & Reim. Achnanthidium deflexum (Reimer) Kingston DMB
BAP ACHDELI Achnanthes delicatula (Kutz.) Grun. Planothidium delicatulum (Kütz.) Round & Bukht. OVB
Psammothidium
BAP ACHDETH Achnanthes detha Hohn & Hellerm. (Hust.) Bukht. & Round OVB
subatomoides
(Cleve) Witkowski, Lange-Bert. &
BAP ACHDISP Achnanthes dispar Cl. Planothidium dispar Metzeltin OVB
BAP ACHDUTH Achnanthes duthii Screen. Achnanthidium duthiei (Sreen.) Edlund OVB
BAP ACHEXIG Achnanthes exigua Grun. Achnanthidium exiguum (Grunow) Czarn. DMB
Achnanthes exigua var.
BAP ACHEXIGC constricta (Grun.) Hust. Achnanthidium exiguum (Grunow) Czarn. DMB
Achnanthes exigua var. Achnanthidium exiguum
BAP ACHEXIGH Krass. (Krasske) Czarn. DMB
heterovalva var. heterovalvum
BAP ACHEXIL Achnanthes exilis Kutz. Achnanthidium exile (Kütz.) Heib. DMB
BAP ACHFLEX Achnanthes flexella (Kutz.) Brun Eucocconeis flexella (Kütz.) F.Meister DMB
Achnanthes flexella var.
BAP ACHFLEXA alpestris Brun Eucocconeis alpestris (Brun) Lange-Bert. OVB
BAP ACHHAUC Achnanthes hauckiana Grun. Planothidium hauckianum (Grunow) Bukht. OVB
Achnanthes hauckiana var.
BAP ACHHAUCR rostrata Schultz DMB
BAP ACHHUNG Achnanthes hungarica (Grun.) Grun. Lemnicola hungarica (Grunow) Round & Basson OVB
BAP ACHKOLB Achnanthes kolbei Hust. Karayevia kolbei (Hust.) Bukht. OVB
BAP ACHKRYO Achnanthes kryophila Pet. Achnanthidium kryophila (J.B.Petersen) Bukht. OVB
Achnanthes kryophila var. Psammothidium
BAP ACHKRYOA africana Choln. subatomoides (Hust.) Bukht. & Round OVB
BAP ACHLANC Achnanthes lanceolata (Breb.) Grun. Planothidium lanceolatum (Bréb. ex Kütz.) Lange-Bert. DMB
Achnanthes lanceolata var.
BAP ACHLANCA abbreviata Reim. Planothidium abbreviatum (Reimer) Potapova OVB
BAP ACHLANCD dubia Grun. Planothidium dubium (Grunow) Round & Bukht. DMB
BAP ACHLANCE elliptica Schulz Planothidium ellipticum (Cleve) Edlund OVB
BAP ACHLANCO omissa Reim. Planothidium joursacense (Hérib.) Lange-Bert. OVB
BAP ACHLANCR rostrata Hust. Planothidium rostratum (Østrup) Lange-Bert. DMB
Achnanthes lapponica var. Eucocconeis lapponica
BAP ACHLAPPN ninckei (Guerm. & Mang.) Reim var. ninckei (Guermeur & Manguin) Edlund DMB
BAP ACHLATE Achnanthes laterostrata Hust. Karayevia laterostrata (Hust.) Bukht. OVB
Psammothidium
BAP ACHLAUE Achnanthes lauenbergiana Hust. lauenburgianum (Hust.) Bukht. & Round OVB
Planothidium
BAP ACHLEMM Achnanthes lemmermanni Hust. (Hust.) E.Morales DMB
lemmermannii
BAP ACHLEVA Achnanthes levanderi Hust. Psammothidium levanderi (Hust.) Bukht. & Round OVB
BAP ACHLINE Achnanthes linearis (W. Sm.) Grun. Rossithidium linearis (W.Sm.) Round & Bukht. DMB
Achnanthidium
BAP ACHLINEC Achnanthes linearis f. curta H.L. Sm. biasolettianum (Grunow) Bukht. DMB
Psammothidium
BAP ACHMARG Achnanthes marginulata Grun. marginulatum (Grunow) Bukht. & Round OVB
Achnanthidium
BAP ACHMICR Achnanthes microcephala (Kutz.) Grun. Kütz. DMB
microcephalum

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Achnanthidium
BAP ACHMINU Achnanthes minutissima Kutz. minutissimum (Kütz.) Czarn. DMB
Achnanthes minutissima Achnanthidium
BAP ACHMINUC Grun. (Grunow) Novais & Van de Vijver DMB
var. cryptocephala neocryptocephalum
BAP ACHOEST Achnanthes oestrupii (Backm. & A. Cl.) Hust. Planothidium oestrupii ( A.Cleve) Edlund OVB
Achnanthes oestrupii var.
BAP ACHOESTL lanceolata Hust. Planothidium pungens (A.Cleve) Lange-Bert. OVB
BAP ACHPINN Achnanthes pinnata Hust. Platessa conspicua (A.Mayer) Lange-Bert. OVB
BAP ACHPLOE Achnanthes ploenensis Hust. Karayevia ploenensis (Hust.) Bukht. OVB
BAP ACHSP Achnanthes sp. OVB
BAP ACHSUBL Achnanthes sublaevis Hust. Psammothidium ventralis (Krasske) Bukht & Round DMB
BAP ACHSUCH Achnanthes suchlandti Hust. Karayevia suchlandtii (Hust.) Bukht. OVB
BAP APLPELL Amphipleura pellucida (Kütz.) Kütz. DMB
BAP APLRUTI Amphipleura rutilans (Trente.) Cl. Berkeleya rutilans (Trentep. ex Roth) Grunow DMB
BAP AMACOFF Amphora coffeiformis (Ag.) Kutz. Halamphora coffeaeformis (C.Agardh) Levkov DMB
BAP AMAHEMI Amphora hemicycla Stoermer & J.J.Yang DMB
BAP AMAINAR Amphora inariensis Krammer DMB
BAP AMANEGL Amphora neglecta Stoermer & J.J.Yang DMB
BAP AMAOVAL Amphora ovalis (Kütz.) Kütz. DMB
BAP AMAOVALA Amphora ovalis var. affinis (Kutz.) V.H. ex DeT. Amphora affinis Kütz. DMB
Amphora ovalis var.
BAP AMAOVALP pediculus (Kutz.) V.H. ex DeT. Amphora pediculus (Kütz.) Grunow DMB
BAP AMAPERP Amphora perpusilla (Grun.) Grun. Halamphora perpusilla (Grunow) Q-M.Wang & Kociolek DMB
BAP AMASP Amphora sp. DMB
BAP AMATENU Amphora tenuistriata Manguin DMB
BAP AMATHUM Amphora thumensis (Mayer) Cl.-Euler. Halamphora thumensis (A.Mayer) Levkov DMB
Amphora veneta var. Halamphora
BAP AMAVENEC capitata Haworth oligotraphenta (Lange-Bert) Levkov DMB
Anomoeoneis serians var.
BAP ANOSERIB brachysira (Breb.) Hust. Brachysira brebissonii DMB
BAP ANOSP Anomoeoneis sp. DMB
BAP ANOVITR Anomoeoneis vitrea (Grun.) Ross Brachysira vitrea (Grunow) R.Ross DMB
BAP ASTFORM Asterionella formosa Hassall RTB
Asterionella formosa var.
BAP ASTFORMG gracillima (Hantzsch) Grunow RTB
BAP ASTRALF Asterionella ralfsii W.Sm. RTB
Caloneis bacillaris var.
BAP CALBACIT (Grun.) A. Cl.(?) Caloneis thermalis (Grunow) Krammer OVB
thermalis
BAP CALBACI Caloneis bacillum (Grunow) Cleve OVB
BAP CALHYAL Caloneis hyalina Hust. OVB
BAP CALSP Caloneis sp. OVB
Caloneis ventricosa var.
BAP CALVENTM (Grunow) Mills OVB
minuta
Caloneis ventricosa var. Caloneis ventricosa var.
BAP CALVENTT truncata Grun. truncatula (Grunow) F.Meister OVB
BAP COCDIMI Cocconeis diminuta Pant. Cocconeis neodiminuta Krammer OVB
BAP COCDISC Cocconeis disculus (Schum.) Cleve OVB
BAP COCPEDI Cocconeis pediculus Ehrenb. OVB
BAP COCPLAC Cocconeis placentula Ehrenb. OVB
Cocconeis placentula var.
BAP COCPLACE euglypta (Ehrenb.) Grunow OVB
BAP COCPLACL lineata (Ehrenb.) Van Heurck OVB
BAP COCPLACR rouxii (Hérib. & Brun) Cleve OVB
BAP COCCSP Cocconeis sp. DBL
BAP COCSPB Cocconeis sp. Ehr. OVB
BAP COCTHUM Cocconeis thumensis A. Mayer Cocconeis neothumensis Krammer OVB
BAP CYMANGUA Cymatopleura angulata Grev. DBB
BAP CYMELLI Cymatopleura elliptica (Bréb. ex Kütz.) W.Sm. OVB
BAP CYMSOLE Cymatopleura solea (Bréb.) W.Sm. DBB
Cymatopleura solea var.
BAP CYMSOLEA apiculata (W.Sm.) Ralfs DBB
BAP CYMSOLER (Ehr.) Grun. Surirella regula Ehrenb. DBB
regula

2019 UPDATED
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(IF APPLICABLE)
BAP CYMSOLES subconstricta D.M. Surirella comperei (Cocquyt & R.Jahn) Cocquyt & R.Jahn DBB
BAP CYMAFFI Cymbella affinis Kütz. DMB
Cymbopleura
BAP CYMAMPH Cymbella amphicephala Nag. amphicephala (Nägeli) Krammer DMB
BAP CYMANGUB Cymbella angustata (W. Sm.) Cl. Cymbopleura angustata (W.Sm.) Krammer DMB
BAP CYMCESA Cymbella cesatii (Rabh.) Grun. ex A.S. Encyonopsis cesatii (Rabenh.) Krammer DMB
BAP CYMCIST Cymbella cistula (Ehr.) Kirchn. Cymbella neocistula Krammer DMB
Cymbella cistula var. Cymbella neocistula var.
BAP CYMCISTG gibbosa Brun islandica Krammer DMB
Cymbella cistula var.
BAP CYMCISTM (Kutz.) V.H.) Cymbella cistula Krammer DMB
maculata
BAP CYMCUSP Cymbella cuspidata Kutz. Cymbopleura apiculata Krammer DMB
BAP CYMCYMB Cymbella cymbiformis C.Agardh DMB
BAP CYMDELI Cymbella delicatula Kutz. Delicata delicatula (Kütz.) Krammer DMB
BAP CYMDILU Cymbella diluviana (Krasske) C. E. Cymbellafalsa diluviana (Krasske) Lange-Bert. & Metzeltin DMB
BAP CYMHUST Cymbella hustedtii Krasske DMB
BAP CYMHYBR Cymbella hybrida Grun. Cymbopleura hybrida (Grunow) Krammer DMB
BAP CYMLAEV Cymbella laevis Nägeli DMB
BAP CYMLANC Cymbella lanceolata (C.Agardh) C.Agardh DMB
Cymbella leptoceros var.
BAP CYMLEPTR rostrata Hust. Cymbella designata Krammer DMB
BAP CYMLUNA Cymbella lunata W. Sm. Encyonema lunatum (W.Sm) Van Heurck DMB
BAP CYMMEXI Cymbella mexicana (Ehrenb.) Cleve DMB
BAP CYMMICR Cymbella microcephala Grun. Encyonopsis microcephala (Grunow) Krammer DMB
BAP CYMMINU Cymbella minuta Hilse ex Rabh. Encyonema minutum (Hilse) D.G.Mann DMB
BAP CYMMINUL Cymbella minuta f. latens (Krasske) Reim. Encyonema latens (Krasske) D.G.Mann DMB
Cymbella minuta var. Encyonema minutum var.
BAP CYMMINUP (Choln.) Reim. (Cholnoky) Czarn. DMB
pseudogracilis pseudogracilis
Cymbella minuta var.
BAP CYMMINUS silesiaca (Bleisch ex Rabh.) Reim. Encyonema silesiacum (Bleisch) D.G.Mann DMB
Cymbopleura
BAP CYMNAVI Cymbella naviculiformis Auersw. (Auersw.) Krammer DMB
naviculiformis
BAP CYMNORV Cymbella norvegica Grun. Encyonema norvegicum (Grunow) A.Mayer DMB
BAP CYMOBTU Cymbella obtusiuscula Kütz. DMB
BAP CYMPROS Cymbella prostrata (Berk.) Cl. Encyonema leibleinii (C.Agardh) Silva et al. DMB
Cymbella prostrata var.
BAP CYMPROAU (Rabh.) Reim. Encyonema auerswaldii Rabenh. DMB
auerswaldii
BAP CYMPUSI Cymbella pusilla Grun. Navicymbula pusilla (Grunow) Krammer 2003 DMB
BAP CYMSINU Cymbella sinuata Greg. Reimeria sinuata (W.Greg.) Kociolek & Stoermer DMB
Cymbella sinuata var.
BAP CYMSINUA (Grun.) Cl. Reimeria sinuata f. antiqua (Grunow) Kociolek & Stoermer DMB
antiqua
BAP CYMSP Cymbella sp. DMB
BAP CYMTRIA Cymbella triangulum (Ehr.) Cl. Encyonema triangulum (Ehrenb.) Kütz. DMB
BAP CYMTUMIA Cymbella tumida (Bréb.) Van Heurck DMB
BAP CYMTUMIU Cymbella tumidula Grunow DMB
BAP DENELEG Denticula elegans Kütz. OVB
BAP DENSP Denticula sp. DMB
BAP DENSUBT Denticula subtilis Grunow DMB
BAP DENTENU Denticula tenuis Kütz. DMB
Denticula tenuis var. (Nägeli ex Kütz.) West &
BAP DENTENUC OVB
crassula G.S.West
BAP DIAANCE Diatoma anceps (Ehr.) Kirchn. Odontidium anceps (Ehrenb.) Ralfs RTB
BAP DIAHIEM Diatoma hiemale (Roth) Heib. Odontidium hyemale (Roth) Kütz. RTB
Diatoma hiemale var.
BAP DIAHIEMM mesodon (Ehr.) Grun. Odontidium mesodon (Ehrenb.) Kütz. RTB
BAP DIASP Diatoma sp. RTB
BAP DIATENU Diatoma tenue Ag. Diatoma tenuis C.Agardh RTB
Diatoma tenue var.
BAP DIATENUE elongatum Lyngb. Diatoma tenuis C.Agardh RTB
BAP DIAVULG Diatoma vulgare Bory. Diatoma vulgaris Bory RTB
BAP DIPBOLD Diploneis boldtiana Cleve OVB
BAP DIPELLI Diploneis elliptica (Kütz.) Cleve OVB
BAP DIPOBLO Diploneis oblongella (Nägeli ex Kütz.) A.Cleve OVB
BAP DIPOCUL Diploneis oculata (Bréb.) Cleve OVB

2019 UPDATED
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BAP DIPOVAL Diploneis ovalis (Hilse) Cleve OVB
BAP DIPPARM Diploneis parma Cleve OVB
BAP DIPPSEU Diploneis pseudovalis (Schum.) Cleve OVB
BAP DIPPUEL Diploneis puella (Schum.) Cl. OVB
BAP DIPSPE Diploneis sp. (J.W.Bailey) Riemer OVB
(J.W. Bail.) Reim. in Patr.
BAP ENTORNA Entomoneis ornata & Reim. OVB
BAP ENTSP Entomoneis sp. OVB
BAP EPISPX Epithemia sp. OVB
BAP EUNCURV Eunotia curvata (Kutz.) Lagerst. Eunotia bilunaris (Ehrenb.) Schaarschm. DMB
BAP EUNINCI Eunotia incisa W.Sm. ex W.Greg DMB
BAP EUNPECT Eunotia pectinalis (Kütz.) Rabenh. DMB
BAP EUNPRAE Eunotia praerupta Ehrenb. DMB
BAP EUNSP Eunotia sp. DMB
Pseudostaurosira
BAP FRABREV Fragilaria brevistriata Grun. brevistriata (Grunow) D.M.Williams & Round DMB
Fragilaria brevistriata var. Pseudostaurosira
BAP FRABREVI inflata (Pant.) Hust. brevistriata var. inflata (Pant.) Edlund DMB
BAP FRACAPU Fragilaria capucina Desm. DMB
Fragilaria capucina var.
BAP FRACAPUL lanceolata Grunow DMB
Fragilaria capucina var.
BAP FRACAPUM (Rabh.) Grun. Fragilaria mesolepta Rabenh. DMB
mesolepta
BAP FRACONS Fragilaria construens (Ehr.) Grun. Staurosira construens Ehrenb. DMB
Fragilaria construens var. Staurosira construens var.
BAP FRACONSB binodis (Ehr.) Grun. binodis (Ehrenb.) Hamilton DMB
Fragilaria construens var. Staurosira construens var. (Temp & H.Perag.) N.A.Andresen,
BAP FRACONSM Temp. & Perag. DMB
minuta minuta Stoermer & Kreis
BAP FRACONSP pumila Grun. pumila (Grunow) Kingston DMB
Fragilaria construens var.
BAP FRACONSS Hust. Staurosira subsalina (Hust.) Lange-Bert. DMB
subsalina
BAP FRACONSV venter (Ehr.) Grun. venter (Ehrenb.) Hamilton DMB
BAP FRACROT Fragilaria crotonensis Kitton DMB
Fragilaria crotonensis var.
BAP FRACROTO oregona Sovereign DMB
BAP FRAINTE Fragilaria intermedia Grun. Fragilaria vaucheriae (Kütz.) J.B.Petersen DMB
Fragilaria intermedia var. (Grunow) Stoermer &
BAP FRAINTEF DMB
fallax J.J.Yang
BAP FRALAPP Fragilaria lapponica Grun. Staurosirella lapponica (Gronow) D.M.Williams & Round OVB
BAP FRALEPT Fragilaria leptostauron (Ehr.) Hust. Staurosirella leptostauron (Ehrenb.) D.M.Williams & Round DMB
Fragilaria leptostauron var. Staurosirella leptostauron
BAP FRALEPTD (Grun.) Hust. (Grunow) Edlund DMB
dubia var. dubia
Fragilariforma
BAP FRANITZ Fragilaria nitzschioides Grun. nitzschioides (Grunow) Lange-Bert. DMB
BAP FRAPINN Fragilaria pinnata Ehr. Staurosirella pinnata (Ehrenb.) D.M.Williams & Round OVB
Fragilaria pinnata var. Staurosirella pinnata var.
BAP FRAPINNI (Grun.) Hust. (Grunow) Hamilton OVB
intercedens intercedens
Fragilaria pinnata var.
BAP FRAPINNL lancettula (Schum.) Hust. Punctastriata lancettula (Schum.) Hamilton & Siver DMB
BAP FRASPC Fragilaria sp. DMB
BAP FRAVAUC Fragilaria vaucheriae (Kutz.) Peters. DMB
Fragilaria vaucheriae var.
BAP FRAVAUCC capitellata (Grun.) Patr. Fragilaria recapitellata Lange-Bert. & Metzeltin DMB
BAP FRAVIRE Fragilaria virescens Ralfs Fragilariforma virescens (Ralfs) D.M.Williams & Round DMB
Frustulia rhomboides var.
BAP FRURHOMA (Grun.) Cl. Frustulia amphipleuroides (Grunow) A.Cleve DMB
amphipleuroides
Frustulia rhomboides var.
BAP FRURHOMS saxonica (Rabh.) DeT. Frustulia saxonica Rabenh. DMB
BAP FRUVULG Frustulia vulgaris (Thwaites) De Toni DMB
BAP GOMACUM Gomphonema acuminatum Ehrenb. OVB
BAP GOMAFFI Gomphonema affine Kütz. OVB
Gomphonema affine var.
BAP GOMAFFII insigne (Greg.) Andrews Gomphonema insigne W.Greg. OVB

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(IF APPLICABLE)
BAP GOMANGU Gomphonema angustatum (Kütz.) Rabenh. OVB
Gomphonema angustatum
BAP GOMANGUP var. productum Grun. Gomphonema productum (Grunow) Lange-Bert. & E.Reichardt OVB
Gomphosphenia (Lange-Bert. & E.Reichardt) Lange-
BAP GOMBRAS Gomphonema brasiliense Grun. lingulatiformis Bert OVB
BAP GOMCLEV Gomphonema clevei Fricke Gomphoneis clevei (Fricke) M.Gil OVB
BAP GOMDICH Gomphonema dichotomum Kütz. OVB
BAP GOMGRAC Gomphonema gracile Ehrenb. OVB
BAP GOMOLIV Gomphonema olivaceum (Hornem.) Bréb. OVB
BAP GOMPARV Gomphonema parvulum (Kütz.) Kütz. OVB
BAP GOMSIMU Gomphonema simus Hohn & Hellerm Gomphonema simum M.H.Hohn & Hellerman OVB
BAP GOMSP Gomphonema sp. OVB
Gomphonema subclavatum
BAP GOMSUBCM (Grunow) R.M.Patrick OVB
var. mexicanum
BAP GOMSUBT Gomphonema subtile Ehrenb. OVB
BAP GOMTENE Gomphonema tenellum Kütz. OVB
BAP GOMTERG Gomphonema tergestinum (Grunow) Fricke OVB
BAP GOMVENT Gomphonema ventricosum W.Greg. OVB
BAP GYRACUM Gyrosigma acuminatum (Kütz.) Rabenh. RTB
BAP GYRATTE Gyrosigma attenuatum (Kütz.) Rabenh. RTB
BAP GYRNODI Gyrosigma nodiferum (Grun.) Reim. Gyrosigma sciotoense (Sull.) Cleve RTB
(W.Sm.) J.W.Griff. &
BAP GYROBSC Gyrosigma obscurum Henfr. RTB
BAP GYRSCAL Gyrosigma scalproides (Rabenh.) Cleve RTB
BAP GYRSCIO Gyrosigma sciotense (Sulliv. & Wormley) Cl. Gyrosigma sciotoense (Sull.) Cleve RTB
BAP GYRSP Gyrosigma sp. RTB
BAP GYRSPEN Gyrosigma spencerii Quek. Gyrosigma acuminatum (Kütz.) Rabenh. RTB
Gyrosigma spencerii var.
BAP GYRSPENC (Grunow) Reimer RTB
curvula
BAP HANARCU Hannaea arcus (Ehrenb.) R.M.Patrick OVB
BAP HANAMPH Hantzschia amphioxys (Ehrenb.) Grunow OVB
Hantzschia amphioxys f.
BAP HANAMPHC capitata O.Müll. OVB
BAP MERCIRC Meridion circulare (Grev.) C.Agardh OVB
Meridion circulare var.
BAP MERCIRCC constrictum (Ralfs) Brun OVB
BAP NAVACCE Navicula acceptata Hust. Geissleria acceptata (Hust.) Lange-Bert. & Metzeltin OVB
BAP NAVANGL Navicula anglica Ralfs Placoneis elginensis (W.Greg.) E.J.Cox OVB
Navicula anglica var.
BAP NAVANGLU subsalsa (Grunow) Cleve OVB
BAP NAVARVE Navicula arvensis Hust. Sellaphora arvensis (Hust.) C.E.Wetzel & Ector OVB
BAP NAVATOM Navicula atomus (Kutz.) Grun. Mayamaea atomus (Kütz.) Lange-Bert OVB
BAP NAVAURO Navicula aurora Sovereign OVB
BAP NAVBACI Navicula bacillum Ehr. Sellaphora bacillum (Ehrenb.) D.G.Mann OVB
(Grunow) Lange-Bert, Metzeltin &
BAP NAVCAPI Navicula capitata Ehr. Hippodonta capitata Witkowski DMB
Navicula capitata var. (Grunow) Lange-Bert, Metzeltin &
BAP NAVCAPIH (Grun.) Ross Hippodonta hungarica DMB
hungarica Witkowski
Navicula capitata var. (Grunow) Lange-Bert, Metzeltin &
BAP NAVCAPIL luneburgensis (Grun.) Patr. Hippodonta luneburgensis Witkowski DMB
BAP NAVCINC Navicula cincta (Ehrenb.) Ralfs OVB
BAP NAVCITR Navicula citrus Krasske Craticula citrus (Krasske) E.Reichardt OVB
BAP NAVCLEM Navicula clementis Grun. Placoneis clementis (Grunow) E.J.Cox OVB
(Gregory ex Grev.) D.G.Mann &
BAP NAVCOCC Navicula cocconeiformis Greg. ex Grev. Cavinula cocconeiformis A.J.Stickle OVB
BAP NAVCONF Navicula confervacea Kutz. Diadesmis confervacea Kütz. OVB
Navicula contenta var. Diadesmis contenta var.
BAP NAVCONTB biceps (Arn.) V.H. biceps (Grunow) P.B.Hamilton OVB
(Grunow) Lange-Bert, Metzeltin &
BAP NAVCOST Navicula costulata Grun. in Cl. & Grun. Hippodonta costulata Witkowski OVB
BAP NAVCRYP Navicula cryptocephala Kütz. DMB
Navicula cryptocephala var.
BAP NAVCRYPV veneta (Kutz.) Rabh. Navicula veneta Kütz. OVB
BAP NAVCUSP Navicula cuspidata (Kutz.) Kutz. Craticula cuspidata (Kütz.) D.G.Mann DMB
BAP NAVDECU Navicula decussis Ostr. Navigeia decussis (Østrup ) Bukht. DMB

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BAP NAVDETE Navicula detenta Hust. OVB
BAP NAVDISJ Navicula disjuncta Hust. (?) Sellaphora disjuncta (Hust.) D.G.Mann OVB
BAP NAVELGIL Navicula elginensis var. lata (M. Perag.) Patr. Placoneis lata (Perag.) R.L.Lowe OVB
BAP NAVEXIG Navicula exigua Greg. ex Grun. Placoneis exigua (W.Greg.) Mereschk. OVB
Navicula exigua var.
BAP NAVEXIGC capitata R.M.Patrick OVB
BAP NAVEXPL Navicula explanata Hust. Placoneis explanata (Hust.) Mayama OVB
BAP NAVFART Navicula farta Hust. OVB
BAP NAVFOSS Navicula fossilis Krasske Mayamaea fossalis (Krasske) Lange-Bert. OVB
BAP NAVFRAC Navicula fracta Hust. Fallacia fracta (Hust. ex Simonsen) D.G.Mann OVB
BAP NAVFRUG Navicula frugalis Hust. Craticula subminuscula (Manguin) C.E.Wetzel & Ector OVB
BAP NAVGAST Navicula gastrum (Ehr.) Kutz. Placoneis gastrum (Ehrenb.) Mereschk. OVB
Navicula gastrum var.
BAP NAVGASTS Hust. OVB
signata
BAP NAVGOTT Navicula gottlandica Grunow OVB
BAP NAVGRACO Navicula graciloides A. Mayer sensu Hust. Navicula cari Ehrenb. DMB
BAP NAVGREG Navicula gregaria Donkin DMB
BAP NAVHAMB Navicula hambergii Hust. Placoneis hambergii (Hust.) Bruder OVB
BAP NAVHARD Navicula harderi Hust. Sellaphora harderi (Hust.) Foets &C.E.Wetzel OVB
Chamaepinnularia
BAP NAVHASS Navicula hassiaca Krasske hassiaca (Krasske) Cantonati & Lange-Bert. OVB
BAP NAVHELE Navicula helensis Schutz Fallacia helensis (P.F.F.Schulz) D.G.Mann OVB
(W.Bock) Levkov, Metzeltin &
BAP NAVIMBR Navicula imbricata Bock Luticola imbricata OVB
A.Pavolv
BAP NAVINGR Navicula ingrata Krasske OVB
BAP NAVINTE Navicula integra (W. Sm.) Ralfs Prestauroneis integra (W.Sm.) Bruder OVB
BAP NAVJAER Navicula jaernefeltii Hust. Cavinula jaernefeltii (Hust.) D.G.Mann & A.J.Stickle OVB
BAP NAVLACU Navicula lacustris Greg. Lacustriella lacustris (W.Greg.) Lange-Bert. & Kulikovskiy OVB
BAP NAVLAEV Navicula laevissima Kutz. Sellaphora laevissima (Kütz.) D.G.Mann OVB
BAP NAVLANC Navicula lanceolata (Ag.) Kutz. Navicula trivialis Lange-Bert. DMB
BAP NAVLATE Navicula latens Krasske Navigeia thingvallae (Østrup) Bukht. OVB
Chamaepinnularia
BAP NAVMEDI Navicula mediocris Krasske mediocris (Krasske) Lange-Bert OVB
BAP NAVMENI Navicula menisculus Schum. DMB
Navicula menisculus var.
BAP NAVMENIU upsaliensis (Grun.) Grun. Navicula upsaliensis (Grunow) Perag. DMB
BAP NAVMINI Navicula minima Grun. OVB
BAP NAVMINU Navicula minuscula Grun. Adlafia minuscula (Grunow) Lange-Bert. OVB
Navicula minuscula var. Adlafia minuscula var.
BAP NAVMINUM muralis (Grun.) Lange-Bert. muralis (Grunow) Lange-Bert. OVB
BAP NAVMURAF Navicula muraliformis Hust. OVB
Adlafia minuscula var.
BAP NAVMURAS Navicula muralis Grun. (Grunow) Lange-Bert. OVB
muralis
BAP NAVMUTI Navicula mutica Kutz. Luticola mutica (Kütz.) D.G.Mann OVB
BAP NAVMUTIC Navicula mutica var. cohnii (Hilse) Grun. Luticola cohnii (Hilse) D.G.Mann OVB
Navicula mutica var.
BAP NAVMUTIU undulata (Hilse) Grun. Luticola undulata (Hilse) D.G.Mann OVB
BAP NAVOCHR Navicula ochridana Hust. OVB
BAP NAVODIO Navicula odiosa J.H.Wallace DMB
BAP NAVOMIS Navicula omissa Hust. Fallacia omissa (Hust.) D.G.Mann OVB
BAP NAVPAUC Navicula paucivisitata R.M.Patrick OVB
BAP NAVPELL Navicula pelliculosa Hilse Fistulifera pelliculosa (Kütz.) Lange-Bert. OVB
(Grunow) R.L.Lowe, Kociolek,
BAP NAVPERP Navicula perpusilla (Kutz.) Grun. Humidophila perpusilla J.R.Johans., Van de Vijver, Lange- OVB
Bert. & Kopalová
BAP NAVPHYL Navicula phyllepta Kütz. OVB
BAP NAVPLAC Navicula placentula (Ehr.) Kutz. Paraplaconeis placentula (Ehrenb.) Kulikovskiy & Lange-Bert. OVB
BAP NAVPLAT Navicula platysoma Ehrenb. OVB
BAP NAVPORI Navicula porifera Hust. Placoneis porifera (Hust.) T.Ohtsuka & Y.Fujita OVB
(Grunow) I.W.Bishop, Minerovic,
BAP NAVPROT Navicula protracta Grun. Prestauroneis protracta Q.Liu & Kociolek OVB
BAP NAVPSEUL Navicula pseudolanceolata Lange-Bert. OVB
BAP NAVPSEUM Navicula pseudomuralis Hust. (?) Fallacia pseudomuralis (Hust.) D.G.Mann OVB
BAP NAVPSEUR Navicula pseudoreinhardtii R.M.Patrick DMB

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Cavinula
BAP NAVPSEUS Navicula pseudoscutiformis Hust. pseudoscutiformis (Hust.) D.G.Mann & Stickle OVB
BAP NAVPSEUY Navicula pseudoventralis Hust. Sellaphora pseudoventralis (Hust.) Chudaev & Gololobova OVB
BAP NAVPUPU Navicula pupula Kutz. Sellaphora pupula (Kütz.) Mereschk. OVB
BAP NAVPUPUE Navicula pupula f. elliptica Hust. Sellaphora wummensis J.R.Johans. OVB
Navicula pupula var.
BAP NAVPUPUC capitata Hust. Sellaphora parapupula Lange-Bert. OVB
BAP NAVPUPUM Navicula pupula var. mutata (Krasske) Hust. Sellaphora mutata (Krasske) Lange-Bert. OVB
BAP NAVRADI Navicula radiosa Kütz. DMB
BAP NAVRADIP Navicula radiosa var. parva Wallace Navicula radiosafallax Lange-Bert. DMB
Navicula radiosa var.
BAP NAVRADIT tenella (Breb.) Cl. & Moll. Navicula cryptotenella Lange-Bert. DMB
BAP NAVREIN Navicula reinhardtii (Grunow) Grunow OVB
Navicula reinhardtii var.
BAP NAVREINE elliptica Hérib. OVB
BAP NAVRHYN Navicula rhynchocephala Kutz. Navicula rhynchotella Lange-Bert. OVB
Navicula rhynchocephala
BAP NAVRHYNA (Kutz.) Grun. Navicula rhynchotella Lange-Bert. OVB
var. amphiceros
BAP NAVSALI Navicula salinarum Grunow OVB
Navicula salinarum var.
BAP NAVSALII intermedia (Grun.) Cl. Navicula capitatoradiata H.Germ. ex Gasse OVB
BAP NAVSAXO Navicula saxophila Bock Luticola saxophila (W.Bock ex Hust.) D.G.Mann OVB
BAP NAVSCHO Navicula schoenfeldii Hust. Geissleria schoenfeldii (Hust.) Lange-Bert. & Metzeltin OVB
BAP NAVSCUT Navicula scutelloides W. Sm. Cavinula scutelloides (W.Sm.) Lange-Bert. OVB
BAP NAVSEMIO Navicula seminuloides Hust. OVB
BAP NAVSEMIU Navicula seminulum Grun. Sellaphora seminulum (Grunow) D.G.Mann OVB
BAP NAVSIMI Navicula similis Krasske emend. Hust. Placogeia similis (Krasske) Bukht. OVB
BAP NAVSP Navicula sp. OVB
BAP NAVSPLE Navicula splendicula VanLand. DMB
BAP NAVSTRO Navicula stroemii Hust. Sellaphora stroemii (Hust.) H.Kobayasi OVB
BAP NAVSUBH Navicula subhamulata Grun. Fallacia subhamulata (Grunow) D.G.Mann OVB
Navicula subhamulata var.
BAP NAVSUBHU Hust. OVB
undulata
BAP NAVSUBMI Navicula submitis Hust. Fallacia submitis (Hust.) D.G.Mann OVB
BAP NAVSUBMU Navicula submuralis Hust. OVB
BAP NAVSUBO Navicula subocculata Hust. OVB
(Hust.) C.E.Wetzel, Ector, Van de
BAP NAVSUBR Navicula subrotundata Hust. Sellaphora subrotundata OVB
Vijver, Compère & D.G.Mann
BAP NAVSUBT Navicula subtilissima Cl. Kobayasiella subtilissima (Cleve) Lange-Bert. OVB
BAP NAVTANT Navicula tantula Hust. Eolimna tantula (Hust.) Lange-Bert. OVB
BAP NAVTENE Navicula tenelloides Hust. OVB
BAP NAVTRIP Navicula tripunctata (O.F.Müll.) Bory OVB
Navicula tripunctata var.
BAP NAVTRIPS schizonemoides (Van Heurck) R.M.Patrick OVB
BAP NAVTRIV Navicula trivialis Lange-Bert. OVB
BAP NAVTUSC Navicula tuscula Ehr. Aneumastus tusculus (Ehrenb.) D.G.Mann & Stickle DMB
BAP NAVTUSCM Navicula tuscula f. minor Hust. Aneumastus minor Lange-Bert OVB
BAP NAVTUSCR Navicula tuscula f. rostrata Hust. Aneumastus rostratus (Hust.) Lange-Bert. 2001 OVB
BAP NAVUTER Navicula utermoehlii Hust. Sellaphora utermoehlii (Hust.) C.E.Wetzel & D.G.Mann OVB
BAP NAVVIRI Navicula viridula (Kütz.) Ehrenb. OVB
Navicula viridula var.
BAP NAVVIRIA avenacea (Bréb.) Van Huerck OVB
Navicula viridula var.
BAP NAVVIRIR rostellata (Kutz.) Cl. Navicula rostellata Kütz. OVB
BAP NAVVITA Navicula vitabunda Hust. Sellaphora vitabunda (Hust.) D.G.Mann OVB
BAP NAVVULP Navicula vulpina Kütz. DMB
BAP NAVWITT Navicula wittrockii (Lagst.) A. Cl.-Eu. Sellaphora laevissima (Kütz.) D.G.Mann OVB
BAP NAVZANO Navicula zanoni Hust. Navicula zanonii Hust. OVB
BAP NEIAFFI Neidium affine (Ehrenb.) Pfitzer OVB
BAP NEIDUBI Neidium dubium (Ehrenb.) Cleve OVB
BAP NEISP Neidium sp. OVB
BAP NITACCO Nitzschia accomodata Hust. DMB
BAP NITACICO Nitzschia acicularioides Hust. Nitzschia spiculum DMB
BAP NITACICS Nitzschia acicularis (Kütz.) W.Sm. DMB

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BAP NITACUL Nitzschia acula (Kütz.) Hantzsch DMB
BAP NITACUM Nitzschia acuminata (W.Sm.) Grun. Tryblionella acuminata W.Sm. DMB
BAP NITACUT Nitzschia acuta Hantzsch DMB
BAP NITALPI Nitzschia alpina Hust. DMB
BAP NITAMPH Nitzschia amphibia Grun. DMB
Nitzschia amphibia var.
BAP NITAMPHA activiscula Grunow DMB
BAP NITANGU Nitzschia angustata (W. Sm.) Grun. Tryblionella angustata W.Sm. DMB
BAP NITANGA Nitzschia angustatula Lange-Bertalot 1987 Tryblionella angustatula (Lange-Bert.) Cantonati & Lange-Bert. DMB
BAP NITAPIC Nitzschia apiculata (Greg.) Grun. Tryblionella apiculata W.Greg. DMB
BAP NITARCH Nitzschia archibaldii Lange-Bert. DMB
BAP NITBACA Nitzschia bacata Hust. DMB
BAP NITCAPI Nitzschia capitellata Hust. DMB
BAP NITCLAU Nitzschia clausii Hantzsch DMB
BAP NITCLOS Nitzschia closterium (Ehr.) W. Sm. Cylindrotheca closterium (Ehrenb.) Reimann & J.C.Lewin DMB
BAP NITCOMM Nitzschia communis Rabenh. DMB
BAP NITCONF Nitzschia confinis Hust. DMB
BAP NITDENT Nitzschia denticula Grun. Denticula kuetzingii Grunow DMB
BAP NITDISS Nitzschia dissipata (Kütz.) Rabenh. DMB
Nitzschia dissipata var.
BAP NITDISSM (Hantzsch) Grunow DMB
media
BAP NITEPIP Nitzschia epiphytica O.Müll. DMB
BAP NITFONT Nitzschia fonticola (Grunow) Grunow DMB
BAP NITFRUS Nitzschia frustulum (Kütz.) Grunow DMB
Nitzschia frustulum var.
BAP NITFRUST Grun. Nitzschia perminuta (Grunow) Perag. DMB
perminuta
Nitzschia frustulum var.
BAP NITFRUSP perpusilla (Rabenh.) Van Heurck DMB
BAP NITFRUT Nitzschia fruticosa Hust. DMB
Nitzschia
BAP NITGAND gandersheimiensis Krasske DMB
BAP NITGRACF Nitzschia graciliformis Lange-Bert. & Simonsen DMB
BAP NITGRACS Nitzschia gracilis Hantzsch DMB
BAP NITHANT Nitzschia hantzschiana Rabenh. DMB
BAP NITHOLL Nitzschia hollerupensis Foged DMB
BAP NITIMPR Nitzschia impressa Hust. DMB
BAP NITINCO Nitzschia inconspicua Grunow DMB
BAP NITINTE Nitzschia intermedia Hantzsch DMB
BAP NITKUETA Nitzschia kuetzingiana Hilse DMB
BAP NITKUETO Nitzschia kuetzingioides Hust. Nitzschia pumila DMB
BAP NITLACU Nitzschia lacuum Lange-Bert. DMB
BAP NITLATE Nitzschia latens Hust. DMB
BAP NITLAUE Nitzschia lauenburgiana Hust. DMB
BAP NITLINE Nitzschia linearis (Agardh) W.Sm. DMB
BAP NITLUZO Nitzschia luzonensis Hust. DMB
BAP NITMEDI Nitzschia mediocris Hust. DMB
BAP NITMICR Nitzschia microcephala Grunow DMB
BAP NITMINUA Nitzschia minuta Bleisch DMB
BAP NITMINUU Nitzschia minutula Grunow DMB
BAP NITOBSI Nitzschia obsidialis Hust. DMB
BAP NITOVAL Nitzschia ovalis H.J.Arn. DMB
BAP NITPALEA Nitzschia palea (Kütz.) W.Sm. DMB
BAP NITPALED Nitzschia palea var. debilis (Kütz.) Grunow DMB
Nitzschia palea var.
BAP NITPALET tenuirostris Grunow DMB
BAP NITPALEC Nitzschia paleacea Grunow DMB
BAP NITPARV Nitzschia parvula Lewis Nitzschia brevissima Grunow DMB
BAP NITPERM Nitzschia perminuta (Grunow) Perag. DMB
BAP NITPSEU Nitzschia pseudofonticola Hust. DMB
BAP NITPUMI Nitzschia pumila Hust. DMB
BAP NITPURA Nitzschia pura Hust. DMB
BAP NITPUSI Nitzschia pusilla Grunow DMB
BAP NITRECT Nitzschia recta Hantzsch ex Rabenh. DMB
BAP NITROMA Nitzschia romana Grun. Nitzschia fonticola (Grunow) Grunow DMB

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BAP NITROST Nitzschia rostellata Hust. DMB
BAP NITSIGM Nitzschia sigmoidea (Nitzsch) W.Sm. DMB
Nitzschia sinuata var.
BAP NITSINUT (Grunow) Grunow DMB
tabellaria
BAP NITSOCI Nitzschia sociabilis Hust. DMB
BAP NITSP Nitzschia sp. DMB
BAP NITSPICO Nitzschia spiculoides Hust. DMB
BAP NITSPICU Nitzschia spiculum Hust. DMB
BAP NITSUBA Nitzschia subacicularis Hust. DMB
BAP NITSUBC Nitzschia subcommunis Hust. DMB
BAP NITSUBL Nitzschia sublinearis Hust. DMB
BAP NITTENU Nitzschia tenuis W.Sm. DMB
BAP NITTHER Nitzschia thermalis (Ehrenb.) Auersw. DMB
BAP NITTROP Nitzschia tropica Hust. Nitzschia fonticola (Grunow) Grunow DMB
Nitzschia tryblionella var.
BAP NITTRYBD debilis (Arn.) A. Mayer Tryblionella debilis H.J.Arn. ex O'Meara OVB
Nitzschia tryblionella var. Tryblionella gracilis var.
BAP NITTRYBS subsalina Grun. subsalina (O'Meara) Aboal OVB
Nitzschia tryblionella var.
BAP NITTRYBV victoriae Grun. Tryblionella victoriae Grunow OVB
BAP NITVALD Nitzschia valdestriata Aleem & Hust. DMB
BAP NITVERM Nitzschia vermicularis (Kütz.) Hantzsch DMB
BAP OPEMART Opephora martyi Herib. Staurosirella martyi (Hérib.) E.Morales & Manoylov OVB
BAP OPESP Opephora sp. OVB
Pinnularia interrupta var.
BAP PININTEM minutissima Hust. Pinnularia subinterrupta Krammer & S.Schroet. OVB
BAP PINLATA Pinnularia lata (Bréb.) Rabenh. OVB
BAP PINMICR Pinnularia microstauron (Ehrenb.) Cleve OVB
BAP PINRUPE Pinnularia rupestris Hantzsch OVB
BAP PINSP Pinnularia sp. OVB
BAP PINSUBC Pinnularia subcapitata W.Greg. OVB
Pinnularia viridis var.
BAP PINVIRIC commutata (Grunow) Cleve OVB
Plagiotropis lepidoptera var.
BAP PLALEPIP proboscidia (Cleve) Reimer DMB
BAP RHOCURV Rhoicosphenia curvata (Kutz.) Grun. ex Rabh. Rhoicosphenia abbreviata (C.Agardh) Lange-Bert. OVB
BAP RHOSPB Rhoicosphenia sp. OVB
BAP STUKRIE Stauroneis kriegeri R.M.Patrick DMB
BAP STUSMIT Stauroneis smithii Grunow DMB
Stauroneis smithii var.
BAP STUSMINC incisa Pant. DMB
Stauroneis smithii var.
BAP STUSMVMI E.Y.Haw. DMB
minima
BAP STUASP Stauroneis sp. DMB
BAP SURANGU Surirella angusta Kütz. OVB
BAP SURBIRO Surirella birostrata Hust. ex Ant.Mayer OVB
Surirella biseriata var.
BAP SURBISEP Meist. OVB
bifrons f. punctata
Surirella linearis var.
BAP SURLINEC constricta Grun. Surirella grunowii Kulikovskiy, Lange-Bert. & Witkowski OVB
BAP SUROVAL Surirella ovalis Bréb. OVB
BAP SUROVAT Surirella ovata Kutz. Surirella brebissonii Krammer & Lange-Bert. OVB
BAP SUROVATP Surirella ovata var pinnata (W. Sm.) Hust. Surirella minuta Bréb. ex Kütz. OVB
BAP SUROVATS Surirella ovata var. salina (W. Sm.) Hust. Surirella salina W.Sm. OVB
BAP SURSP Surirella sp. W.Sm. OVB
BAP SURTURG Surirella turgida W. Sm. OVB
BAP SYNACUS Synedra acus Kutz. Ulnaria acus (Kütz.) Aboal DMB
Fragilaria
BAP SYNAMPH Synedra amphicephala Kutz. amphicephaloides Lange-Bert DMB
Synedra amphicephala var.
BAP SYNAMPHA austriaca (Grun.) Hust. Fragilaria austriaca (Grunow) Lange-Bert. DMB
BAP SYNCYCL Synedra cyclopum Brutschy DMB
BAP SYNDELI Synedra delicatissima W. Sm. Ulnaria delicatissima (W.Sm.) Aboal & P.C.Silva DMB
Synedra delicatissima var. Ulnaria delicatissima var.
BAP SYNDELIA angustissima Grun. angustissima (Grunow) Aboal & P.C.Silva DMB

2019 UPDATED
(IF APPLICABLE)
(IF APPLICABLE)
BAP SYNDEME Synedra demerarae Grun. Fragilaria demerarae (Grunow) Lange-Bert. DMB
BAP SYNFAMI Synedra familica Kütz. DMB
BAP SYNFILI Synedra filiformis Grunow DMB
Synedra filiformis var.
BAP SYNFILIE A.Cleve DMB
exilis
BAP SYNMINU Synedra miniscula Grun. Synedra famelica (Kütz.) Lange-Bert DMB
BAP SYNNANA Synedra nana F.Miester DMB
BAP SYNOSTE Synedra ostenfeldii (Krieg.) A.Cleve DMB
Pseudostaurosira
BAP SYNPARA Synedra parasitica W. Sm. (W.Sm.) E.Morales DMB
parasitica
Synedra parasitica var. Pseudostaurosira
BAP SYNPARAS subconstricta (Grun.) Grun. subconstricta (Grunow) Kulikovskiy & Genkal DMB
(Ralfs ex Kütz.) D.M.Williams &
BAP SYNPULC Synedra pulchella Ralfs ex Kutz Ctenophora pulchella Round DMB
BAP SYNRADI Synedra radians Kutz. Fragilaria radians (Kütz.) D.M.Williams & Round DMB
BAP SYNRUMP Synedra rumpens Kutz. Fragilaria rumpens (Kütz.) G.W.F.Carlson DMB
Synedra rumpens var.
BAP SYNRUMPS Grunow DMB
scotica
BAP SYNSOC Synedra socia Wallace DMB
BAP SYNSPP Synedra sp. DMB
BAP SYNSP3 Synedra sp. # 3 in house taxon DMB
BAP SYNTENE Synedra tenera W. Sm. Fragilaria tenera (W.Sm.) Lange-Bert. DMB
BAP SYNULNA Synedra ulna (Nitz.) Ehr. Ulnaria ulna (Nitzsch) Compère DMB
BAP SYNULNAB Synedra ulna var. biceps Kutz. Ulnaria biceps (Kütz) Compère DMB
BAP SYNULNAH Synedra ulna var. chaseana B.W.Thomas DMB
BAP SYNULNAO Synedra ulna var. contracta Venkt. Ulnaria contracta (Østrup) E.Morales and M.L.Vis DMB
BAP SYNULNAD Synedra ulna var. danica (Kutz.) V.H. Ulnaria danica (Kütz) Compère & Bukht. DMB
Synedra ulna var.
BAP SYNULNAL (W. Sm.) Brun. Ulnaria biceps (Kütz) Compère DMB
longissima
BAP TABFENE Tabellaria fenestrata (Lyngb.) Kütz. RTB
BAP TABFLOC Tabellaria flocculosa (Roth) Kütz. RTB
Tabellaria flocculosa var.
BAP TABFLOCG (A.Cleve) B.M.Knudson TFG
geniculata
BAP TABSP Tabellaria sp. RTB
BAP UNIPENN Unidentified Pennales DMB
CHL ACASP Acanthosphaera sp. OVO
Actinastrum aciculare f.
CHL ACTACICM (Hub.-Pest.) Compère OVO
minimum
CHL ACTGRAC Actinastrum gracilimum G.M.Sm. OVO
CHL ACTHANT Actinastrum hantzschii Lagerh. OVO
CHL ANKBRAU Ankistrodesmus braunii (Naeg.) Brun. Chlorolobion braunii (Nägeli) Komárek FUS
Ankistrodesmus convolutus
CHL ANKCONVM (Nägeli) Rabenh. FUS
var. minutus
CHL ANKFALC Ankistrodesmus falcatus (Corda) Ralfs FUS
Ankistrodesmus falcatus
CHL ANKFALCF var. fasciculatus Margalef FUS
Ankistrodesmus falcatus (W. & G.S. West) G.S.
CHL ANKFALCM var. mirabilis West Monoraphidium mirabile (West & G.S.West) Pankow FUS
CHL ANKGELI Ankistrodesmus gelifactum (Chod.) Bourr. Elakatothrix gelifacta (Chodat) Hindák FUS
CHL ANKGRAC Ankistrodesmus gracilis (Reins.) Kors. Messastrum gracile (Reinsch) T.S.Garcia FUS
CHL ANKSETI Ankistrodesmus setigerus (Schroed.) G.S. West Schroederia setigera (Schröd.) Lemmerm. FUS
CHL ANKSPI Ankistrodesmus sp. FUS
CHL ANKSPIR Ankistrodesmus spiralis (W.B.Turner) Lemmerm. FUS
CHL ANKSTIP Ankistrodesmus stipitatus Komárk.-Legn. FUS
CHL AKYJUDA Ankyra judayi (G.M.Sm.) Fott FUS
CHL AKYLANC Ankyra lanceolata (Kors.) Fott Lanceola spatulifera (Korshikov) Hindák FUS
CHL AKYSP Ankyra sp. FUS
CHL ARTBIFI Arthrodesmus bifidus Breb. Octacanthium bifidum (Bréb.) Compère OVO
CHL ARTSP Arthrodesmus sp. OVO
CHL ARTTRIA Arthrodesmus triangularis Lag. Staurodesmus triangularis (Lagerh.) Teiling OVO
CHL ASTSUPE Asterococcus superbus (Cienk.) Scherff. OVO
CHL BOTBRAU Botryococcus braunii Kütz. OVO
CHL BOTSPC Botryococcus sp. OVO
CHL BOTSPS Botryosphaera sp. OVO

2019 UPDATED
(IF APPLICABLE)
(IF APPLICABLE)
CHL CARCORD Carteria cordiformis (H.J.Carter) Diesing OVO
CHL CARSP Carteria sp. OVO
CHL CARWISC Carteria wisconsinensis Hub.-Pest. OVO
CHL CHMSP Characium sp. FUS
CHL CHLGLOB Chlamydomonas globosa J.Snow OVO
CHL CHLSP Chlamydomonas sp. OVO
CHL CRLSP Chlorella sp. OVO
CHL CBHSP Chlorobrachis sp. OVO
CHL CHGMINI Chlorogonium minimum Playfair OVO
CHL CHGSP Chlorogonium sp. OVO
CHL CDASUBS Chodatella subsalsa Lemm. Lagerheimia subsala Lemmerm. OVO
CHL CDPSP Chodatellopsis sp. OVO
(Chodat/G.M.Sm)
CHL CLOACICC Closteriopsis acicularis J.H.Belcher & Swale FUS
CHL CLOLONG Closteriopsis longissima (Lemmerm.) Lemmerm. FUS
Closteriopsis longissima
CHL CLOLONGA var. acicularis G.M. Sm. Closteriopsis acicularis (Chodat/G.M.Sm) J.H.Belcher & Swale FUS
CHL CLOSPC Closteriopsis sp. FUS
CHL CLOACICD Closterium aciculare T.West FUS
Closterium aciculare var.
CHL CLOACICS subpronum W. & G.S. West Closterium aciculare T.West FUS
Closterium acutum var.
CHL CLOACUTV variabile (Lemmerm.) Willi Krieg. FUS
CHL CLOEXIL Closterium exile West & G.S.West FUS
CHL CLOGRAC Closterium gracile Bréb. ex Ralfs FUS
CHL CLOPARV Closterium parvulum Nägeli FUS
CHL CLOSPD Closterium sp. FUS
Closterium strigosum var.
CHL CLOSTRIE (G.S.West) Willi Krieg. FUS
elegans
CHL COCBICE Coccoid bicells DBL
CHL COCFUSI Coccoid fusiform FUS
CHL COCOVAL Coccoid oval OVO
CHL COCSP4 Coccoid sp. #4 OVO
CHL COCSPHE Coccoid sphere OVO
CHL COEASTR Coelastrum astroideum De Not. OVO
CHL COECAMB Coelastrum cambricum W.Archer OVO
CHL COEMICR Coelastrum microporum Nägeli OVO
CHL COEMORU Coelastrum morus W. & G.S. West Coelastrum sphaericum Nägeli OVO
Coelastrum
CHL COEPSEU Korshikov OVO
pseudomicroporum
CHL COERETI Coelastrum reticulatum (Dang.) Senn. Hariotina reticulata P.A.Dang. OVO
CHL COESPT Coelastrum sp. OVO
CHL COESPHA Coelastrum sphaericum Nägeli OVO
CHL COESPY Coenocystis sp. OVO
CHL CORSP? Coronastrum sp. (?) OVO
CHL COSBOTR Cosmarium botrytis Menegh. ex Ralfs OVO
CHL COSDEPR Cosmarium depressum (Nägeli) P.Lundell OVO
CHL COSMELA Cosmarium melanosporum W.Archer & J.Roy OVO
CHL COSPHAS Cosmarium phaseolus Bréb. ex Ralfs OVO
CHL COSREGN Cosmarium regnellii Wille OVO
CHL COSSPD Cosmarium sp. OVO
CHL COSSUBC Cosmarium subcostatum Nordst. OVO
CHL CRUFENE Crucigenia fenestrata (Schmidle) Schmidle OVO
CHL CRUIRRE Crucigenia irregularis Wille Crucigeniella irregularis (Wille) P.M.Tsarenko & D.M.John OVO
CHL CRUPULC Crucigenia pulchra (W. & G. S. West) Kom. Crucigeniella apiculata (Lemmerm.) Komárek OVO
CHL CRUQUAD Crucigenia quadrata Morren OVO
(A.Braun) D.M.John, M.J.Wynne &
CHL CRURECT Crucigenia rectangularis A. Braun Willea rectangularis P.M.Tsarenko OVO
CHL CRUSP Crucigenia sp. OVO
CHL CRUTETR Crucigenia tetrapedia (Kirchn.) Kuntze OVO
(G.M.Sm.) D.M.John, M.J.Wynne &
CHL CRUTRUN Crucigenia truncata G.M. Sm. Willea truncata OVO
P.M.Tsarenko
CHL CROCRAS Crucigloea crassiseta (Skuja) Soeder OVO
CHL DACINFU Dactylococcus infusionum Nägeli FUS

2019 UPDATED
(IF APPLICABLE)
(IF APPLICABLE)
Dictyosphaerium
CHL DICEHRE ehrenbergianum Nägeli OVO
(H.Bachm.) Krienitz, C.Bock,
CHL DICELEG Dictyosphaerium elegans (Bachm.) Mychonastes elegans OVO
Dadheech & Pröschold
Dictyosphaerium Mucidosphaerium (H.C.Wood) C.Bock, Pröschold,
CHL DICPULC pulchellum Wood pulchellum Krientz OVO
CHL DICSP Dictyosphaerium sp. OVO
CHL DIDANOM Didymogenes anomala (G.M.Sm.) Hindák FUS
CHL DIDPALA Didymogenes palatina Schmidle FUS
CHL DIDSPG Didymogense sp. OVO
CHL DIMLUNA Dimorphococcus lunatus A.Braun OVO
CHL DIMSP Dimorphococcus sp. OVO
CHL DIPDECU Diplochloris decussata Korshikov FUS
CHL DIPLUNA Diplochloris lunata (Fott) Fott FUS
CHL ECHSPC Echinocoleum sp. OVO
CHL ECHLIMN Echinosphaerella limnetica G.M.Sm. OVO
CHL ECHSPS Echinosphaeridium sp. OVO
CHL ELAGENE Elakatothrix genevensis (Reverdin) Hindák FUS
CHL ELASP Elakatothrix sp. FUS
CHL ELAVIRI Elakatothrix viridis (Snow) Printz Fusola viridis J.Snow FUS
CHL ENACOEL Enallax coelastroides (Bohl.) Skuja OVO
CHL EUDELEG Eudorina elegans Ehrenb. OVO
CHL EUDSP Eudorina sp. OVO
CHL EUTSPC Eutetramorus sp. OVO
CHL FRADROE Franceia droescheri (Lemmerm.) G.M.Sm. OVO
CHL FRAELON Franceia elongata Korshikov OVO
CHL FRAMINU Franceia minuscula Hind. OVO
CHL FRAOVAL Franceia ovalis (Francé) Lemmerm. OVO
CHL FRASPB Franceia sp. OVO
Dichotomococcus
CHL GLOLIMN Gloeactinium limneticum G.M. Sm. Korshikov OVO
curvatus
CHL GLOBACI Gloeocystis bacillus (Teil.) Fott Chlamydocapsa bacillus (Teiling) Fott OVO
Chlamydocapsa
CHL GLOGIGA Gloeocystis gigas (Kutz.) Lag. planctonica (West & G.S.West) Fott OVO
Chlamydocapsa
CHL GLOPLAN Gloeocystis planktonica (W. & G.S. West) Lemm. planctonica (West & G.S.West) Fott OVO
CHL GLOSPC Gloeocystis sp. OVO
CHL GLOSPT Gloeotila sp. CYL_TUBE
CHL GOLMAXI Golenkinia maxima Tiffany & Ahlstrom OVO
CHL GOLRADI Golenkinia radiata Chodat OVO
Golenkinia radiata var.
CHL GOLRADIB brevispina Tiff. & Ahlstr. Golenkinia brevispina Korshikov OVO
CHL GOLSPK Golenkinia sp. OVO
CHL GOLSPI Golenkiniopsis sp. OVO
CHL GONPECT Gonium pectorale O.F.Müll. OVO
CHL GONSP Gonium sp. OVO
CHL GYRCORD Gyromitus cordiformis Skuja CON
CHL HETGALL Heterodesmus gallicus Bourr. & Coute FUS
CHL KIRCONT Kirchneriella contorta (Schm.) Bohlin Raphidocelis danubiana (Hindák) Marvan, Komárek & Comas FUS
Pseudokirchneriella
CHL KIRELON Kirchneriella elongata G.M. Sm. elongata (G.M.Sm.) Hindák FUS
CHL KIRLUNA Kirchneriella lunaris (Kirchn.) K.Möbius FUS
(G. S. West) Kom.-Legn. (G.S.West) Marvan, Komárek &
CHL KIRMAYO Kirchneriella mayori Raphidocelis mayorii FUS
in Kom. Comas
CHL KIROBES Kirchneriella obesa (West) West & G.S.West FUS
Kirchneriella obesa var.
CHL KIROBESM major (Bern.) G.M. Sm. Kirchneriella major C.Bernard FUS
CHL KIRSP Kirchneriella sp. FUS
Nephrochlamys
CHL KIRSUBS Kirchneriella subsolitaria G. S. West subsolitaria (G.S.West) Korshikov FUS
CHL KORLIMN Korshikoviella limnetica (Lemmerm.) P.C.Silva FUS
CHL LAGBALA Lagerheimia balatonica (Scherff.) Hindák OVO
CHL LAGCHOD Lagerheimia chodatii C.Bernard OVO
CHL LAGCILI Lagerheimia ciliata (Lagerh.) Chodat OVO

2019 UPDATED
(IF APPLICABLE)
(IF APPLICABLE)
CHL LAGCING Lagerheimia cingula G.M. Sm. Lagerheimia circumfilata (Seligo) E.Hegewald & A.Schmidt OVO
CHL LAGCITR Lagerheimia citriformis (J.Snow) Collins OVO
Lagerheimia citriformis var.
CHL LAGCITRP Tiffany & Ahlstrom OVO
paucispina
CHL LAGGENE Lagerheimia genevensis (Chodat) Chodat OVO
CHL LAGLONG Lagerheimia longiseta (Lemmerm.) Printz OVO
Lagerheimia longiseta var.
CHL LAGLONGM major G.M.Sm. OVO
CHL LAGQUAD Lagerheimia quadriseta (Lemmerm.) G.M.Sm. OVO
CHL LAGSPL Lagerheimia sp. OVO
CHL LAGSUBS Lagerheimia subsalsa Lemmerm. OVO
CHL LAGWRAT Lagerheimia wratislaviensis Schröd. OVO
CHL LOBSPC Lobocystis sp. OVO
Lobomonas ampla var.
CHL LOBAMPLO okensis Korschikov OVO
CHL LOBSPM Lobomonas sp. OVO
Micractinium Micractinium
CHL MITBORN (Cour.)Kors. (W.Conrad) Korshikov OVO
bornheimiense bornhemiense
CHL MITPUSI Micractinium pusillum Fresen. OVO
CHL MITQUAD Micractinium quadrisetum (Lemmerm.) G.M.Sm. OVO
CHL MITSP Micractinium sp. Fresnius OVO
CHL MPASPA Microspora sp. CYL_TUBE
CHL MONARCU Monoraphidium arcuatum Kors. Ankistrodesmus arcuatus Korshikov FUS
(Nag. in Kutz.) Kom.-
CHL MONBRAU Monoraphidium braunii Legn. Chlorolobion braunii (Nägeli) Komárek FUS
CHL MONCIRC Monoraphidium circinale (Nygaard) Nygaard FUS
CHL MONCONT Monoraphidium contortum (Thur.) Komárk.-Legn. FUS
Monoraphidium
CHL MONCONV (Corda) Komárk.-Legn. FUS
convolutum
(Wolosz.) Hindák &
CHL MONDYBO Monoraphidium dybowskii Komárk.-Legn. FUS
CHL MONGRIF Monoraphidium griffithii (Berk.) Komárk.-Legn. FUS
CHL MONIRRE Monoraphidium irregulare (G.M.Sm.) Komárk.-Legn. FUS
CHL MONMINU Monoraphidium minutum (Nägeli) Komárek.-Legn. FUS
(Korshikov) Komárk.-
CHL MONOBTU Monoraphidium obtusum Legn. FUS
CHL MONPUSI Monoraphidium pusillum (Printz) Komárk.-Legn. FUS
CHL MONSAXA Monoraphidium saxatile Komárk.-Legn. FUS
Monoraphidium
CHL MONSETI Monoraphidium setiforme (Nyg.) Kom.-Legn. komarkovae Nygaard FUS
CHL MONSKUJ Monoraphidium skujae Fott FUS
CHL MONSPH Monoraphidium sp. FUS
(West & G.S.West)
CHL MONTORT Monoraphidium tortile Komárk.-Legn. FUS
CHL MOUSP Mougeotia sp. CYL_TUBE
CHL NEODANU Neodesmus danubialis Hindák OVO
CHL NCHPSP Nephrochlamys sp. FUS
CHL NCHSUBS Nephrochlamys subsolitaria (G.S.West) Korshikov FUS
CHL NCHWILL Nephrochlamys willeana (Printz) Korshikov FUS
CHL NCTAGAR Nephrocytium agardhianum Nägeli FUS
Nephrocytium W. West in W. & G.S.
CHL NCTECDY Oonephris obesa (West & G.S.West) Fott FUS
ecdysiscepanum West.
CHL NCTLIMN Nephrocytium limneticum (G.M.Sm.) G.M.Sm. FUS
CHL NCTSP Nephrocytium sp. FUS
CHL OEDSP Oedogonium sp. CYL_TUBE
CHL OOCBORG Oocystis borgei J.Snow OVO
CHL OOCCRAS Oocystis crassa Witt. Neglectella solitaria (Wittr.) Stenclová & Kaštovský OVO
CHL OOCELLIP Oocystis elliptica West OVO
CHL OOCELVMI Oocystis elliptica f. minor W. West Oocystis elliptica West OVO
Oocystis gigas var.
CHL OOCGIVIN incrassata West & G.S.West OVO
CHL OOCLACU Oocystis lacustris Chodat OVO
CHL OOCMARS Oocystis marssonii Lemmerm. OVO
CHL OOCNATAM Oocystis natans v. major G.M. Smith Oocystis natans var. major G.M.Sm. OVO

2019 UPDATED
(IF APPLICABLE)
(IF APPLICABLE)
CHL OOCNODU Oocystis nodulosa West & G.S.West OVO
CHL OOCPARV Oocystis parva West & G.S.West OVO
CHL OOCPUSI Oocystis pusilla Hansg. OVO
CHL OOCRHOM Oocystis rhomboidea Fott OVO
CHL OOCSOLI Oocystis solitaria Witt. Neglectella solitaria (Wittr.) Stenclová & Kaštovský OVO
CHL OOCSP Oocystis sp. OVO
CHL OOCSUBM Oocystis submarina Lagerh. OVO
CHL PANMORU Pandorina morum (O.F.Müll.) Bory OVO
CHL PANSP Pandorina sp. OVO
CHL PARMULT Paradoxia multiseta Svirenko CLA
CHL PAUTENE Paulschulzia tenera (Korshikov) J.W.G.Lund OVO
CHL PEDBIRA Pediastrum biradiatum Meyer 1829 Parapediastrum biradiatum (Meyen) E.Hegewald OVB
Pseudopediastrum
CHL PEDBORY Pediastrum boryanum (Turp.) Menegh. boryanum (Turpin) E.Hegwald OVB
CHL PEDDUPL Pediastrum duplex Meyen OVB
Pediastrum duplex var.
CHL PEDDUPLC clathratum (A. Braun) Lag. Pediastrum duplex Meyen OVB
CHL PEDDUPLG W. & G.S. West Lacunastrum gracillimum (West & G.S.West) H.McManus OVB
gracillimum
CHL PEDDUPLR reticulatum Lag. Pediastrum duplex Meyen OVB
Pediastrum integrum var.
CHL PEDINTEP Printz (?) Stauridium privum (Printz) E.Hegewald OVB
priva
CHL PEDSIMP Pediastrum simplex (Meyen) Lemm. Monactinus simplex (Meyen) Corda OVB
Monactinus simplex var.
CHL PEDSIMPE Pediastrum simplex v. echin Wittrock echinulatum (Wittr.) M.C.Pérez, Maidana & Comas OVB
Pediastrum simplex var.
CHL PEDSIMPD (Bail.) Rabh. Monactinus simplex (Meyen) Corda OVB
duodenarium
CHL PEDSP Pediastrum sp. OVB
CHL PEDTETR Pediastrum tetras (Ehr.) Ralfs Stauridium tetras (Ehrenb.) E.Hegewald OVB
Pediastrum tetras var.
CHL PEDTETRT (Corda) Hansg. OVB
tetraedon
CHL PHAMINU Phacotus minuscula Bourr. Phacotus minusculus Bourr. OVO
CHL PHASPC Phacotus sp. OVO
CHL PHYSP Phythelios sp. OVO
CHL PLKLAUT Planktonema lauterborni Schm. Binuclearia lauterbornii (Schmidle) Proshk.-Lavr. CYL_TUBE
CHL PLKSP Planktonema sp. Planctonema sp. CYL_TUBE
CHL PKAGELA Planktosphaeria gelatinosa G.M.Sm. OVO
CHL PTESP Pteromonas sp. OVO
CHL PYRSP Pyramidomonas sp. CON
CHL QUACHOD Quadrigula chodatii (Tanner-Füll.) G.M.Sm. FUS
CHL QUACLOS Quadrigula closterioides (Bohlin) Printz OVO
CHL QUALACU Quadrigula lacustris (Chod.) G.M. Sm. Gregiochloris lacustris (Chodat) Marvan, Komárek & Comas OVO
CHL RAPSIGM Raphidiocelis sigmoidea Hindák FUS
CHL RAYHEMI Rayssiella hemisphaerica Edelst. & Prescott OVO
CHL SCEABUN Scenedesmus abundans (Kirch.) Chod. Desmodesmus abundans (Kirchn.) E.Hegewald OVO
CHL SCEACUM Scenedesmus acuminatus (Lag.) Chod. Tetradesmus lagerheimii M.J.Wynne & Guiry FUS
Scenedesmus acuminatus Pectinodesmus pectinatus
CHL SCEACUMT (Skuja) Uherk. f. tortuosus (Skuja) E.Hegewald FUS
var. tortuosus
CHL SCEACUT Scenedesmus acutus Meyen Tetradesmus obliquus (Turpin) M.J.Wynne FUS
CHL SCEARCU Scenedesmus arcuatus (Lemmerm.) Lemmerm. FUS
CHL SCEARMA Scenedesmus armatus (Chodat) Chodat OVO
Scenedesmus armatus var. Desmodesmus armatus
CHL SCEARMAB (Gugl.-Prinz) Chod. (Guglielm.) E.Hegewald OVO
bicaudatus var. bicaudatus
CHL SCEBALA Scenedesmus balantonicus Hort. Scenedesmus balatonicus Hortob. OVO
CHL SCEBICA Scenedesmus bicaudatus (Hansg.) Chodat OVO
Scenedesmus bicaudatus
CHL SCEBICAB Hortob. OVO
var. brevicaudatus
CHL SCEBIJU Scenedesmus bijuga (Turp.) Lag. Scenedesmus bijugus (Turpin) Lagerh. OVO
CHL SCEBRAS Scenedesmus brasiliensis Bohlin Desmodesmus brasiliensis (Bohlin) E.Hegewald OVO
CHL SCEBREV Scenedesmus brevispina (G.M.Sm.) Chodat OVO
Desmodesmus opoliensis
CHL SCECARI Scenedesmus carinatus (Lemm.) Chod. (Lemmerm.) E.Hegewald OVO
var. carinatus
Desmodesmus
CHL SCEDENT Scenedesmus denticulatus Kirch. denticulatus (Lagerh.) S.S.An, Friedl & E.Hegewald OVO

2019 UPDATED
(IF APPLICABLE)
(IF APPLICABLE)
Scenedesmus denticulatus Desmodesmus
CHL SCEDENTL var. linearis Hansg. denticulatus var. linearis (Hansg.) E.Hegewald OVO
CHL SCEDIMO Scenedesmus dimorphus (Turp.) Kutz. Tetradesmus dimorphus (Turpin) M.J.Wynne OVO
CHL SCEDISP Scenedesmus dispar Breb. Desmodesmus dispar (Bréb.) E.Hegewald OVO
CHL SCEECOR Scenedesmus ecornis (Ehrenb.) Chodat OVO
Scenedesmus ecornis var. Scenedesmus obtusus f.
CHL SCEECORD disciformis Chod. disciformis (Chodat) Compère OVO
Scenedesmus ecornis var.
CHL SCEECORS Hort. & Nemeth OVO
disciformis f. spinosus
CHL SCEELLI Scenedesmus ellipsoideus Chod. Desmodesmus armatus (Chodat) E.Hegewald OVO
CHL SCEGRAN Scenedesmus granulatus W. & G.S. West Desmodesmus granulatus (West & G.S.West) P.M.Tsarenko OVO
CHL SCEINTE Scenedesmus intermedius Chod. Desmodesmus intermedius (Chodat) E.Hegewald OVO
Scenedesmus intermedius
CHL SCEINTEA var. acaudatus Hortob. OVO
Scenedesmus intermedius Desmodesmus intermedius
CHL SCEINTEB var. balatonicus Hort. var. balatonicus (Hortob.) P.M.Tsarenko OVO
Scenedesmus intermedius Desmodesmus intermedius
CHL SCEINTEI var. bicaudatus Hort. var. acutispinus (Y.V.Roll) E.Hegewald OVO
CHL SCEMICR Scenedesmus microspina Chod. Desmodesmus microspina (Chodat) P.M.Tsarenko OVO
CHL SCEOPOL Scenedesmus opoliensis P. Richt. Desmodesmus opoliensis (P.G.Richt.) E.Hegewald OVO
CHL SCEQUAD Scenedesmus quadricauda (Turpin) Bréb. OVO
Scenedesmus quadricauda
CHL SCEQUADB var. biornatus Kiss OVO
Scenedesmus quadricauda Desmodesmus armatus
CHL SCEQUADL var. longispina (Chod.) G.M. Sm var. longispina (Chodat) E.Hegewald OVO
CHL SCEQUADM var. maxima W. & G.S. West Desmodesmus maximus (West & G.S.West) E. Hegewald OVO
CHL SCEQUADQ var. quadrispina (Chod.) G.M. Sm. Desmodesmus abundans (Kirchn.) E.Hegewald OVO
CHL SCEQUADC var.longispina f capricornus (Skuja) Uher. OVO
CHL SCESECU Scenedesmus securiformis Playfair OVO
CHL SCESERR Scenedesmus serratus (Corda) Bohl. Desmodesmus serratus (Corda) S.S.An, Friedl & E.Hegewald OVO
CHL SCESP Scenedesmus sp. OVO
CHL SCESPIC Scenedesmus spicatus W.& G.S. West Desmodesmus subspicatus (Chodat) E.Hegewald & Ant.Schmidt OVO
CHL SCESPIN Scenedesmus spinosus Chod. Desmodesmus spinosus (Chodat) E.Hegewald OVO
CHL SCESUBS Scenedesmus subspicatus Chod. Desmodesmus subspicatus (Chodat) E.Hegewald & Ant.Schmidt OVO
CHL SCEVELI Scenedesmus velitaris Komárek OVO
CHL SCHCOMP Schizoclamys compacta Prescott OVO
Pseudoschroederia
CHL SCHANTI Schroederia antillarum Kom. antillarum (Komárek) E.Hegewald & Schnepf FUS
CHL SCHINDI Schroederia indica Philipose FUS
CHL SCHJUDA Schroederia judayi Ankyra judayi (G.M.Sm.) Fott FUS
CHL SCHSETI Schroederia setigera (Schröd.) Lemmerm. FUS
CHL SLCELLI Sphaerellocystis ellipsoidea H.Ettl OVO
CHL SLCLATE Sphaerellocystis lateralis Fott & Nováková OVO
CHL SPLELON Sphaerellopsis elongata Skvortsov OVO
CHL SPLSP Sphaerellopsis sp. OVO
CHL SPYSCHR Sphaerocystis schroeteri Chodat OVO
CHL STAANAT Staurastrum anatinum Cooke & Wills STR
Staurastrum ellipticum var.
CHL STAELLIM minor West STR
CHL STAGRAC Staurastrum gracile Ralfs ex Ralfs STR
CHL STALACU Staurastrum lacustris G.M. Sm. Staurastrum lacustre G.M.Sm. STR
CHL STALEPT Staurastrum leptocladum Nordst. STR
CHL STAPARA Staurastrum paradoxum Meyen ex Ralfs STR
CHL STASPD Staurastrum sp. STR
CHL STSMAMI Staurodesmus mamillatus (Nordst.) Teiling STR
CHL STISP Stichococcus sp. OVO
CHL TETALTE Tetrachlorella alternans (G.M.Sm.) Korshikov OVO
Tetraedron Tetraëdron
CHL TETARTH (G.S. West) Woloz. Wolosz. CRU
arthrodesmiforme arthrodesmiforme
CHL TETCAUD Tetraedron caudatum (Corda) Hansg. Tetraëdron caudatum (Corda) Hansg. CRU
Tetraedron caudatum var. Tetraëdron caudatum var.
CHL TETCAUDL Lemm. Lemmerm. CRU
longispinum longispinum

2019 UPDATED
(IF APPLICABLE)
(IF APPLICABLE)
CHL TETDUOS Tetraedron duospinum Ackley Tetraëdron duospinum Ackley CRU
Pseudostaurastrum
CHL TETLIMN Tetraedron limneticum Borge limneticum (Borge) Couté & Rousselin CRU
CHL TETLUNU Tetraedron lunula (Reins.) Wille Tetraëdron lunula (Reinsch) Hansg. CRU
CHL TETMINI Tetraedron minimum (A. Braun) Hansg. Tetraëdron minimum (A.Braun) Hansg. CRU
Tetraedron minimum var. Tetraëdron minimum var.
CHL TETMIVTE tetralobulatum Reins tetralobulatum Reinsch CRU
CHL TETMUTI Tetraedron muticum (A. Braun) Hansg. Goniochloris mutica (A.Braun) Fott CRU
CHL TETREGU Tetraedron regulare Kutz. Tetraëdriella regularis (Kütz.) Fott CRU
Tetraedron regulare var.
CHL TETREGUI incus Teil. Chlorotetraëdron incus (Teiling) Komárek & Kováčik CRU
CHL TETTRIG Tetraedron trigonum (Nag.) Hangs. Tetraëdron trigonum (Nägeli) Hansg. CRU
CHL TETREGUL Tetraedron tumidulum (Reins.) Hansg. Tetraëdron tumidulum (Reinsch) Hansg. CRU
Pseudosphaerocystis
CHL TETLACU Tetraspora lacustris Lemm. lacustris (Lemmerm.) Nováková OVO
CHL TETLEMM Tetraspora lemmermannii Fott OVO
(Y.V.Roll) Ahlstrom &
CHL TETSP Tetraspora sp. OVO
Tiffany
CHL TETGLAB Tetrastrum glabrum (Roll) Ahlstr. & Tiff. OVO
CHL TETHETE Tetrastrum heteracanthum (Nordst.) Chodat OVO
Tetrastrum heteracanthum f.
CHL TETHETEE (Playf.) Ahlstr. & Tiff. Tetrastrum elegans Playfair OVO
elegans
Tetrastrum Tetrastrum
CHL TETSTAU staurogeniaeformae (Schroed.) Lemm. staurogeniiforme (Schröd.) Lemmerm. CON
CHL TREPLAN Treubaria planktonica (G.M. Sm.) Kor. Treubaria planctonica (G.M.Sm.) Korshikov OVO
CHL TREQUAD Treubaria quadrispina (G.M.Sm.) Fott & Kováčik OVO
CHL TRESCHM Treubaria schmidlei (Schröd.) Fott & Kováčik CON
CHL TRESETIA Treubaria setigera (W.Archer) G.M.Sm. CON
CHL TRESETIU Treubaria setigerum (Arch.) G.M. Sm. Treubaria setigera (W.Archer) G.M.Sm. CON
CHL TRESP Treubaria sp. CON
CHL TRETRIA Treubaria triappendiculata Bern. Treubaria triappendiculata C.Bernard CON
CHL TROSP Trochiscia sp. OVO
CHL ULOSP Ulothrix sp. CYL_TUBE
Unidentified Colonial
CHL UNICGR greens OVO
Unidentified filamentous
CHL UNIFILA CYL_TUBE
green
Unidentified green
CHL UNICLFLA flagellate OVO
CHL WESSP Westella sp. OVO
CHL XANCONC Xanthidium concinnum Arch. (?) STR
CHM BODSP Bodopsis sp. OVO
CHM VACSP Vacuolaria sp. OVO
CHR BITCHOD Bitrichia chodatii (Reverdin) Chodat OVO
CHR BITLONG Bitrichia longispina (J.W.G.Lund) Bourr. OVO
CHR BITOHRI Bitrichia ohridiana (Fott) Nich. OVO
CHR BITOLLU Bitrichia ollula (Fott) Fott OVO
CHR BITSP Bitrichia sp. OVO
CHR CHMSPM Chromulina sp. OVO
CHR CRNINSI Chrysarachnion insidians Pascher OVO
CHR CHRSP Chrysococcus sp. OVO
CHR CYKANGU Chrysolykos angulatus (Willén) Nauwerck OVO
CHR CYKPLAN Chrysolykos planktonicus Mack. Chrysolykos planctonicus B.Mack OVO
CHR CYKSKUJ Chrysolykos skujae (Nauw.) Bourr. Chrysoikos skujae (Nauwerck) Willén OVO
CHR CYKSP Chrysolykos sp. OVO
CHR CSLCONR Chrysosphaerella conradi Bourr. Chrysosphaerella conradii Bourr. OVO
CHR CSLLONG Chrysosphaerella longispina Lauterborn OVO
CHR CSLRODH Chrysosphaerella rodhei Skuja OVO
CHR CHYSP Chrysosphaerella sp. OVO
CHR CODINCL Codonoeca inclinata (Kent) Kent OVO
CHR DESBRAC Desmarella brachycalyx Skuja OVO
CHR DESMONI Desmarella moniliformis Kent OVO
CHR DESSP Desmarella sp. OVO
CHR DIDSPC Didymochrysis sp. OVO

2019 UPDATED
(IF APPLICABLE)
(IF APPLICABLE)
CHR DINACUM Dinobryon acuminatum Ruttner OVO
CHR DINBALT Dinobryon balticum (Schütt) Lemmerm. OVO
CHR DINBAVA Dinobryon bavaricum O.E.Imhof OVO
Dinobryon bavaricum var.
CHR DINBAVAM (Lemmerm.) Krieg. OVO
medium
Dinobryon bavaricum var.
CHR DINBAVAV vanhoeffenii (Bachm.) Krieg. Dinobryon vanhoeffenii H.Bachm. OVO
CHR DINBORG Dinobryon borgei Lemmerm. OVO
CHR DINCALC Dinobryon calciformis Bachm. Dinobryon caliciforme H.Bachm. OVO
Dinobryon condensatum Dinobryon condensatum
CHR DINCONDP var. planktonicum Skuja var. planctonicum Skuja OVO
CHR DINCREN Dinobryon crenulatum West & G.S.West OVO
CHR DINCYLI Dinobryon cylindricum O.E.Imhof OVO
Dinobryon cylindricum var.
CHR DINCYLIA alpinum (O.E.Imhof) H.Bachm. OVO
Dinobryon cylindricum var.
CHR DINCYLIP Lemmerm. OVO
palustre
CHR DINDIVE Dinobryon divergens O.E.Imhof OVO
Dinobryon divergens var.
CHR DINDIVES schauinslandii (Lemmerm.) Brunnth. OVO
CHR DINEURY Dinobryon eurystoma (A.Stokes) Lemmerm. OVO
CHR DINSERT Dinobryon sertularia Ehrenb. OVO
Dinobryon sertularia var.
CHR DINSERTP protuberans (Lemm.) Kreig. Dinobryon protuberans Lemmerm. OVO
CHR DINSOCI Dinobryon sociale (Ehrenb.) Ehrenb. OVO
Dinobryon sociale var.
CHR DINSOCIA (Brunnth.) H.Bachm. OVO
americanum
Dinobryon sociale var.
CHR DINSOCIS stipitatum (F.Stein) Lemmerm. OVO
CHR DINSP Dinobryon sp. OVO
CHR DINSTOK Dinobryon stokesii Lemm. Epipyxis stokesii (Lemmerm.) G.M.Sm. OVO
Dinobryon stokesii var.
CHR DINSTOKE epiplanktonicum Skuja Epipyxis epiplanctica (Skuja) D.K.Hilliard & Asmund OVO
CHR DINTUBA Dinobryon tubaeformae Nyg. Dinobryon tubaeforme Nygaard OVO
CHR DINUTRI Dinobryon utriculus Stein Epipyxis utriculus (Ehrenb.) Ehrenb. OVO
Dinobryon utriculus var. Epipyxis utriculus var.
CHR DINUTRIA acutum Shil. acuta (J.Schiller) D.K.Hilliard & Asmund OVO
Dinobryon utriculus var.
CHR DINUTRIT Lemm. Epipyxis tabellariae (Lemmerm.) G.M.Sm. OVO
tabellariae
CHR DIPSPI Diplosiga sp. OVO
CHR EPISP Epipyxis sp. OVO
CHR HAPSP Haptophyceae OVO
CHR HYASP Hyalobryon sp. OVO
CHR KEPASPE Kephyrion asper (Lackey) Bourr. OVO
CHR KEPBORE Kephyrion boreale Skuja OVO
CHR KEPCINC Kephyrion cinctum (Lackey) Bourr. OVO
CHR KEPCUPU Kephyrion cupuliformae Conr. Kephyrion cupuliforme W.Conrad OVO
CHR KEPCYLI Kephyrion cylindricum (Lackey) W.Conrad OVO
CHR KEPDOLI Kephyrion doliolum W.Conrad OVO
CHR KEPHEMI Kephyrion hemisphaericum (Lackey) W.Conrad OVO
CHR KEPHILL Kephyrion hilliardii N.H.Nocholls OVO
CHR KEPLITT Kephyrion littorale J.W.G.Lund OVO
CHR KEPMAST Kephyrion mastigophorum G.W.Schmidt OVO
CHR KEPOVAL Kephyrion ovale (Lackey) Hub.-Pest. OVO
CHR KEPPRIS Kephyrion prismaticum W.Conrad OVO
CHR KEPRUBR Kephyrion rubri-claustri W.Conrad OVO
Kephyrion rubri-claustri
CHR KEPRUBRA var. amphora (Lackey) W.Conrad OVO
CHR KEPSP Kephyrion sp. OVO
CHR KEPSPIR Kephyrion spirale (Lackey) W.Conrad OVO
CHR LAGSPR Lagynion sp. OVO
CHR MALACAR Mallomonas acaroides Perty (?) Mallomonas ploesslii Perty OVO
CHR MALAKRO Mallomonas akrokomos Ruttner OVO
CHR MALALLO Mallomonas allorgei (Deflandre) W.Conrad OVO
CHR MALCAUD Mallomonas caudata Iwanoff OVO

2019 UPDATED
(IF APPLICABLE)
(IF APPLICABLE)
CHR MALMAJO Mallomonas majorensis Skuja OVO
Mallomonas
CHR MALPSEU pseudocoronata Prescott OVO
CHR MALRADI Mallomonas radiata W.Conrad OVO
CHR MALSP Mallomonas sp. OVO
CHR MALSP3 Mallomonas sp. #3 OVO
Mallomonas tonsurata var.
CHR MALTONSA alpina (Pasch. & Rutt.) Krieg. (?) Mallamonas alpina Pascher & Ruttner OVO
CHR MALVALK Mallomonas valkanoviana W.Conrad OVO
CHR MASSP Mastigella sp. OVO
CHR MONOVAT Monosiga ovata Kent OVO
CHR MONSPO Monosiga sp. OVO
Ochromonadaceae colonial
CHR OCHCOLO OVO
form
CHR OCHSPOV Ochromonas sp. - ovoid OVO
CHR PARSP Paraphysomonas sp. OVO
CHR PORSP Porochrysis sp. Pascher OVO
CHR PSEACUT Pseudokephyrion acutum J.Schiller OVO
Pseudokephyrion
CHR PSEALAS alaskanum D.K.Hilliard OVO
Pseudokephyrion
CHR PSEATTE attenuatum D.K.Hilliard OVO
CHR PSECONI Pseudokephyrion conicum J.Schiller OVO
Pseudokephyrion Pseudokephyrion
CHR PSECYLI cylindrcum Bourr. cylindricum Bourr. OVO
CHR PSEELLI (Pasch.) Schm. (Pascher) W.Conrad OVO
ellipsoidium ellipsoideum
CHR PSEENTZ Pseudokephyrion entzii W.Conrad OVO
Pseudokephyrion
CHR PSEFORM formosissimum W.Conrad OVO
CHR PSELATU Pseudokephyrion latum (J.Shiller) G.W.Schmidt OVO
CHR PSEMILL Pseudokephyrion millerense K.H.Nicholls OVO
Pseudokephyrion
CHR PSEMINU minutissimum W.Conrad OVO
CHR PSESP Pseudokephyrion sp. OVO
CHR PSEUNDU Scherf. Scherff. OVO
undulatissimium undulatissimum
CHR RHIMAJO Rhizochrysis major Naumann OVO
CHR RHISPC Rhizochrysis sp. OVO
CHR SPISP Spiniferomonas sp. OVO
CHR STCSP Stichogloea sp. OVO
CHR STYAURE Stylotheca aurea (Bachm.) Boloch. Stylochrysalis aurea (Chodat) H.Bachm. OVO
CHR STYSP1 Stylotheca sp. #1 OVO
CHR STYSP?) Stylotheca sp. (?) OVO
CHR SYNSPR Synura sp. OVO
Unidentified chrysophyte
CHR UNCFLAG OVO
flagellate
Unidentified chrysophyte
CHR UNCOVO ovoid (nonflagellate) OVO
CHR UROLIND Uroglena lindii Bourr. Uroglena lindiae Bourr. OVO
CHR UROSP Uroglena sp. OVO
CHR UROVOLV Uroglena volvox Ehrenb. OVO
CRY CRYEROS Cryptomonas erosa Ehrenb. OVO
CRY CRYPHAS Cryptomonas phaseolus Skuja OVO
CRY CRYPYRE Cryptomonas pyrenoidifera Geitler OVO
CRY CRYREFL Cryptomonas reflexa Skuja OVO
CRY CRYROST Cryptomonas rostratiformis Skuja Cryptomonas curvata Ehrenb. OVO
CRY UNICRYP Cryptomonas sp. OVO
CRY RHOLENS Rhodomonas lens Pascher & Ruttner TRP
Plagioselmis (Skuja) Novarino, I.A.N.Lucas &
CRY RHOMINU Rhodomonas minuta Skuja nannoplanctica S.Morrall TRP
CRY RHOSPC Rhodomonas sp. TRP
CRY SENPARV Sennia parvula Skuja OVO
CRY UNICRYP Unidentified cryptomonad OVO

2019 UPDATED
(IF APPLICABLE)
(IF APPLICABLE)
(Rabenh. ex Bornet & Flahault)
CYA ANBCIRC Anabaena circinalis Rabh. Dolichospermum circinale Wacklin, L.Hoffm. & Komárek OVO
Dolichospermum (Bréb. Ex Bornet & Flahualt) Wacklin,
CYA ANBFLOS Anabaena flos-aquae (Lyngb.) Breb. OVO
flosaquae L.Hoffm. & Komárek
CYA ANBSP Anabaena sp. OVO
(Klebhan) Wacklin, L.Hoffm. &
CYA ANBSPIR Anabaena spiroides Kleb. Dolichospermum spiroides Komárek OVO
Anabaena spiroides var. (Klebhan) Wacklin, L.Hoffm. &
CYA ANBSPCR crassa Lemm. Dolichospermum spiroides Komárek OVO
CYA ANACYAN Anacystis cyanea Dr. & Daily Microcystis aeruginosa (Kütz.) Kütz. OVO
CYA ANAMONT Anacystis montana Dr. & Daily Sorospora montana (Lightf.) Hassall OVO
CYA APHFLOS Aphanizomenon flos-aquae (Lyngb.) Ralfs Aphanizomenon flosaquae Ralfs ex Bornet & Flahault CYL_TUBE
CYA APHSP Aphanizomenon sp. CYL_TUBE
CYA APASP Aphanocapsa sp. OVO
CYA APOSP Aphanothece sp. OVO
CYA CRODISP Chroococcus dispersus (Keissl.) Lemmerm. OVO
(Lemmerm.) Komárková, Jezberová,
CYA CROLIMN Chroococcus limneticus Lemm. Limnococcus limneticus OVO
O.Komárek & Zapomelová
CYA CROSP Chroococcus sp. OVO
CYA CROTURG Chroococcus turgidus (Kütz.) Nägeli OVO
CYA COEDUBI Coelosphaerium dubium Grunow OVO
Coelosphaerium
CYA COENAEG Ung. Woronichinia naegeliana (Unger) Elenkin OVO
naegelianum
CYA COESPP Coelosphaerium sp. OVO
CYA CYASP Cyanarcus sp. FUS
CYA DACSP Dactylococcopsis sp. FUS
CYA GLCSP Gloeocapsa sp. OVO
CYA GLTSP Gloeothece sp. OVO
CYA GMAAPON Gomphosphaeria aponina Kütz. OVO
CYA GMALACU Gomphosphaeria lacustris Chod. Snowella lacustris (Chodat) Komárek & Hindák OVO
CYA GMASP Gomphosphaeria sp. OVO
(G.M.Sm.) Komárek, Zapomelová,
CYA LYNBIRG Lyngbya birgei G.M. Sm. Limnoraphis birgei J.Šmarda, Kopecký, Rejmánková, CYL_TUBE
Woodhouse, Neilan & Komárková
(Gomont ex Gomont) Anagn. &
CYA LYNLAGE Lyngbya lagerheimii (Moeb.) Gom. Leptolyngbya lagerheimii Komárek CYL_TUBE
(Lemmerm.) Komárk.-Legn. &
CYA LYNLIMN Lyngbya limneticum Lemm. Planktolyngbya limnetica Cronberg CYL_TUBE
CYA LYNSP1 Lyngbya sp. #1 CYL_TUBE
CYA LYNSPIR Lyngbya spirulinoides Gomont ex Gomont CYL_TUBE
CYA MERSP Merismopedia sp. OVO
CYA MERTENU Merismopedia tenuissima Lemmerm. OVO
CYA MICAERU Microcystis aeruginosa (Kütz.) Kütz. OVO
CYA MICSPA Microcystis sp. OVO
CYA OSCAGAR Oscillatoria agardhii Gom. Planktothrix agardhii (Gomont) Anagn. & Komárek CYL_TUBE
(Gomont) Strunecký, Komárek &
CYA OSCAMOE Oscillatoria amoena (Kutz.) Gom. Microcoleus amoenus CYL_TUBE
J.R.Johans.
CYA OSCBORN Oscillatoria bornetii Zukal Tychonema bornetii (Zukal) Anagn. & Komárek CYL_TUBE
(Bory ex Gomont) Strunecký, Komárek
CYA OSCFORM Oscillatoria formosa Bory Kamptonema formosum & J.Šmarda CYL_TUBE
CYA OSCLIMN Oscillatoria limnetica Lemm. Pseudanabaena limnetica (Lemmerm.) Komárek CYL_TUBE
CYA OSCMINI Oscillatoria minima Gick. Jaaginema minimum (Gicklhorn) Anagn. & Komárek CYL_TUBE
CYA OSCPROL Oscillatoria prolifica (Grev.) Gom. Planktothrix prolifica (Gomont) Anagn. & Komárek CYL_TUBE
CYA OSCRUBE Oscillatoria rubescens De Cand. Planktothrix rubescens (DC. ex Gomont) Anagn. & Komárek CYL_TUBE
CYA OSCSP Oscillatoria sp. CYL_TUBE
CYA OSCSUBB Oscillatoria subbrevis Schmidle CYL_TUBE
CYA OSCTENU Oscillatoria tenuis C.Agardh ex Gomont CYL_TUBE
Oscillatoria tenuis var.
CYA OSCTENUN natans Gom. Lyngbya natans Hansg. CYL_TUBE
Oscillatoria tenuis var. (Rabenh.ex Gomont) Anagn. &
CYA OSCTENUT (Kutz.) Rabh. Phormidium tergestinum CYL_TUBE
tergestina Komárek
CYA PHOINUN Phormidium inundatum Kütz. ex Gomont CYL_TUBE
CYA PHOSP Phormidium sp. CYL_TUBE
CYA RAPSP Raphidiopsis sp. FUS
CYA RHASP Rhabdoderma sp. FUS

2019 UPDATED
(IF APPLICABLE)
(IF APPLICABLE)
CYA SCHSP Schizothrix sp. CYL_TUBE
CYA SPUMAJO Spirulina major Kütz. ex Gomont CYL_TUBE
CYA SPUSP Spirulina sp. CYL_TUBE
CYA SPUSUBT Spirulina subtilissima Kütz. ex Gomont CYL_TUBE
CYA SYESP Synechococcus sp. OVO
Unidentified coccoid
CYA UNICOCCY cyanophyta OVO
Unidentified moniliform
CYA UNIMONIC OVO
coccoid
EUG EUGSP Euglena sp. OVO
EUG EUTSPO Eutreptia sp. OVO
EUG PHASPE Phacus sp. OVO
EUG TRAABRU Trachelomonas abrupta Svirenko OVO
EUG TRAGRAN Trachelomonas granulosa Playfair OVO
EUG TRAHISP Trachelomonas hispida (Perty) F.Stein OVO
EUG TRALACU Trachelomonas lacustris Drezep. OVO
EUG TRASCAB Trachelomonas scabra Playfair OVO
EUG TRASP Trachelomonas sp. OVO
PYR AMPLUTE Amphidinium luteum Skuja OVO
PYR AMPSPP Amphidinium sp. OVO
PYR AMPWIGR Amphidinium wigrense Wolosz. OVO
PYR CERHIRU Ceratium hirundinella (O.F.Müll.) Dujard. CER
PYR GLESP Glenodinium sp. OVO
PYR GYMEXCA Gymnodinium excavatum Nygaard OVO
PYR GYMHELV Gymnodinium helveticum Pen. Gyrodinium helveticum (Penard) Y.Takano & T.Horig. OVO
Gymnodinium helveticum f.
PYR GYMHELVA achroum Skuja OVO
PYR GYMPARA Gymnodinium paradoxum A.J.Schill. OVO
PYR GYMSP Gymnodinium sp. OVO
PYR HEMNASU Hemidinium nasutum F.Stein OVO
PYR HEMSP Hemidinium sp. OVO
PYR PERSP Peridinium sp. OVO
UNI UNICOCCS spherical OVO
UNI UNIFLAG5 Unidentified flagellate #5 OVO
Unidentified flagellate
UNI UNIFLAGO ovoid OVO
Unidentified flagellates
UNI UNIFLAGF FUS
fusiform
UNI UNIHEAR Unidentified heartshaped OVO
UNI UNILORIP Unidentified loricate sp. OVO
UNI UNILORIH Unidentified loricate sphere OVO
XAN CENBELA Centritractus belanophorus (Schmidle) Lemmerm. OVO
XAN CLBPOLY Chlorobotrys polychloris Pascher OVO
XAN CLBSP Chlorobotrys sp. OVO
XAN ISTTRIS Isthmochloron trispinatum (West & G.S.West) Skuja OVO
XAN TETSMIT Tetraplektron smithii (Bourr.) Bourr. (?) OVO
XAN UNICOCCX xanthophyte OVO

Appendix 3: Formulas for Calculating Biomass for Various Phytoplankton Shapes
SHAPE CODE BIOVOLUME FORMULA

“ARC” (3.1416 * width ^ 2 * length) / 12
“BUT” ((4 / 3) * 3.1416 * (width / 2) ^ 2 * (length / 2))
(4 * (1 / 3) * 3.1416 * (diam / 2) ^ 2 * (length)) + (3.1416 * (width / 2) ^ 2 *
“CER”
(depth))
“CLA” ((1 / 3) * 3.1416 * (length) * (width / 2) ^ 2)
“CLB” (3.1416 * (length / 2) * (width / 2) * (depth))
“CON” ((1 / 3) * 3.1416 * (width / 2) ^ 2 * (length))
“CRU” (length ^ 3) / 4
“CYB” (width * depth * (length / 2))
“CYL_DISK” (3.1416 * (diam / 2) ^ 2 * (depth)); Typically observed in “valve view”
(3.1416 * (width / 2) ^ 2 * (length)); Typically observed lying on its side
“CYL_TUBE”
(“girdle view”)
“CYM” (3.1416 * (width / 2) ^ 2 * (length)) / 2
“DBL” ((8 / 3) * 3.1416 * (length / 2) * (width / 2) * (depth / 2))
“DBB” ((8 / 3) * 3.1416 * (length / 2) * (width / 2) * (depth / 2))
“DMB” (width * depth * (length / 2))
“FUS” ((2 / 3) * 3.1416 * (width / 2) ^ 2 * (length / 2))
“LEN” ((2 / 3) * 3.1416 * (width / 2) ^ 2 * (length / 2))
“LUN” ((2 / 3) * 3.1416 * (width / 2) ^ 2 * (length))
“OVB” (3.1416 * (length / 2) * (width / 2) * (depth))
(4 / 3) * 3.1416 * (width / 2) ^ 2 * (length / 2); Diameter values from
“OVO”
spherical specimens, if present, are converted to length and width
“PYR” ((1 / 3) * 3.1416 * (width / 2) ^ 2 * (length))
“RNF” (3.1416 * (width / 2) ^ 2 * (length))
“ROD” (3.1416 * (width / 2) ^ 2 * (length))
“RTB” (length * width * depth)
“SGB” (length * width * depth)
“STR” 6 * ((1 / 3) * 3.1416 * (width / 2) ^ 2 * length)
“TFG” (3.1416 * (width / 2) ^ 2 * depth) + (2 * (diam * length * width))
“TRP” (3.1416 * width ^ 2 * ((length + width) / 2)) / 12

lg401.v07 Phytoplankton Analysis - Rfa

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Standard Operating Procedure

for Phytoplankton Analysis

Version 07, March 2021

1.0 SCOPE AND APPLICATION ............................................................................................................ 1

LG401, Version 07, March 2021 ii

Standard Operating Procedure for Phytoplankton Analysis

2.0 SUMMARY OF METHOD

3.0 SAMPLE COLLECTION

4.2 Compound microscope with an objective system of magnification of 1000x or greater

4.3 Sedimentation chambers: 5-, 10-, 25-, 50- and 100-cc

LG401, Version 07, March 2021 Page 1 of 35

4.4 Beakers: 300- and 600-mL

4.5 Large hotplate capable of boiling water

4.7 Centrifuge tubes, graduated 50-mL

4.8 Coverslips, round, #1 thickness, 22-mm diameter

4.9 Pre-cleaned microscope slides, 25 x 75 mm

4.11 Long-neck disposable pipettes or macropipetter

4.12 Rubber bulbs for pipettes (if disposable pipettes used)

4.13 10-mL autopipette

5.1 HNO3 = Nitric Acid (concentrated)

5.2 H2O2 = Hydrogen peroxide (30% solution)

5.3 K2Cr2O7 = Potassium dichromate

5.4 HyraxTM mounting media, or equivalent (Naphrax, Pleurax, etc.)

5.6 Commercial formaldehyde solution 37 - 40% (= formalin)

5.7 Immersion oil for upright microscopy

6.0 ANALYTICAL PROCEDURE

6.2 10-mL Preliminary Investigation

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10-mL preliminary counts

1) 101 organisms total

Count needed (minimum)

1) 250 organisms total

1) 25-mL sample for sedimentation

The definition of an organism is as follows:

6.3 Sample Sedimentation

LG401, Version 07, March 2021 Page 3 of 35

6.3.3 Approximate settling times necessary are as follows:

100 mL ----------------- 100 hours

6.4 Total Sample Analysis

6.5 Sedimented Sample Analysis

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6.6 Diatom Sample Analysis

6.6.2 Cleaning of Diatoms

Sample cleaning and slide preparation must be performed in the hood.

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6.6.3 Diatom Slide Preparation

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6.6.4 Diatom Enumeration and Identification

6.6.4.1 Diatoms should be identified and enumerated at 1000-1250x. Identification should

6.6.5 General Analysis Guidelines

LG401, Version 07, March 2021 Page 7 of 35

correspond only to previously established taxa. If an analyst encounters a species that

7.1 Soft algae

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7.1.9 Store the archived sample in a pre-labeled tray/box.

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9.0 QUALITY CONTROL AUDITS AND METHOD PRECISION

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Appendix 1: Geometric Forms and the Measurements