C Notes
C Notes
C Notes
The term Horticulture is derived from two Latin words i.e. Hortus meaning garden or enclosure
and Cultra meaning cultivation. So, horticulture literally means garden culture or culture of garden.
Modern horticulture may be defined as a part of agricultural science, which deals with the production,
utilization, and improvement of fruits, vegetables, flowers, ornamentals, plantation crops, medicinal and
aromatic plants etc.
DIVISIONS OF HORTICULTURE
Horticulture crops include fruits, Vegetables, flowers, plantation crops, Spices, condiments,
Medicinal and Aromatic crops etc. In addition to these, Horticulture also deals with raising of trees for
shade, ornamental and avenue purposes, planning and raising of ornamental gardens, parks and raising of
seed and planting material. Further, horticulture also deals with the utilization of horticulture produce and
improvement of horticulture crops.
Hence, based upon the crops dealt and also their purpose and utilization, the branch of horticulture
is sub-divided in to the following divisions for convenience.
Pomology:It is derived from two words i.e. Pomum meaning fruit and Logos meaning discourse or
study. So, pomology is study or cultivation of fruit crops. E.g. Mango, Sapota, Guava, Grape, Banana
etc.
Fruit: It is a developed and matured ovary with or without accessory parts and which is generally
eaten as raw.
Olericulture: It is derived from two words ie. Oleris meaning Potherb and Cultra meaning cultivation.
So, Olericulture literally means potherb cultivation. In the present days it is broadly used to indicate
the cultivation of vegetables. Eg. Brinjal, Okra, Tomato, Pumpkin etc.
Vegetable: It is any part of the herbaceous plant that is generally used after cooking as a principal part
of the meal.
Floriculture: It is derived from two words i.e. Florus meaning flower and Cultra meaning cultivation.
So floriculture means study of flower crops.
In this there are again two sub-divisions. (1) Commercial Floriculture (2) Ornamental Floriculture.
Commercial floriculture:Deals with the cultivation of flower crops grown on commercial scale
for profit (Income). E.g.: Rose, Jasmine, Carnation, Aster, and Marigold etc.
Ornamental floriculture:It deals with the raising of flower crops for ornamental, pleasure and
fashion purposes. E.g.: Dahlia, Zinnia, Cosmos, Hibiscus, Balsam, Nerium, Poinsettia, Hollyhock,
Gerbera, and Gaillardia etc.
Arboriculture: This branch deals with the raising of perennial trees meant for shade, avenue or
ornamental purposes. Eg.Polyalthia, Spathodea, Cassia, Gulmohar etc.
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Fundamentals of Horticulture
Plantation: Are those crops, which are cultivated in an extensive scale in large
areas, owned and managed by an individual or a company and whose produce is utilized only
processin
Eg.Coffee, Tea, Rubber, Coconut, Cocoa
Spices and This branch deals with the cultivation of crops whose produce is used
mainly for seasoning and flavouring
Are those plants the products of which are
Spices s food
madeadjuncts
use to add aroma
flavour.
Eg.Pepper, Cardamom, Clove, Cinnamon, All spice
Condiment:Are those plants the products of which are made use of as food adjuncts to add
only.E .Turmeric,Ginger,Red chillies,Onion,Garlic
Both spices and condiments contain essential oils,
aroma,
which
flavour andand they
are of little nutritive
Medicinal and aromatic : It deals with the cultivation of medicinal plants, which
drugs and aromatic crops
yields aromatic (essential)
Medicinal -are those plants, which are rich in secondary metabolites and are potential
of drugs. The secondary metabolites include alkaloids, glycosides, coumarins, flavonoides and
steroids
Eg. Periwikle, Opium, Menthi, Cinchona, Dioscorea Yam, Belladona, Senna, Sarpagandha,
Aswagandha, Tulasi
Aromatic - are those plants, which possesses essential oils in them. The essential oils are
odoriferous steam volatile constituentsants.
of aromatic
E . Lemon grass, Citronella,
Vetiver, Geranium, Davanam, Lavendor
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Fundamentals of Horticulture
Vitamins: These are the important constituents of fruits and are indispensable part of human diet.
Although required in very minute quantities, they are absolutely essential for the maintenance of health.
The deficiency of any vitamin from the diet for considerable period may lead to diseased state or disorder
conditions. Fruits supply several vitamins
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Fundamentals of Horticulture
1. Per Unit Area Yield is High: As compared to the field crops per hectare yield of horticulture crops
is very high. From an fruit area of land more yield is obtained e.g. paddy gives a maximum yield
of only 30 q/ha, while Banana gives 300 to 500 q/ha, Pine apple 450 q/ha and Grapes 90 - 150 q/ha. In
present shortage of food and scarcity of land by growing fruits more food can be produced.
2. High Returns per Unit Area: From one unit area of land more income will be obtained e.g. Well
kept orchard of apple, grapes and sweet orange can give as much as Rs. 25,0000 per ha as net income.
3. A Free Grower/Labour Remains Engaged for the Whole: An opportunity for maintaining
labours throughout the year like the cereals where one cannot keep himself and employ the labours
during the slack season.
4. Best Utilization of Waste Land:
Some fruit crops can offer best utilization of waste land crops like wood apple, custard apple, karonda,
litchi etc. can be grown in such areas.
5. Food energy: To meet the annual calories requirements of food per year one would have to
cultivate about 0.44 ha of wheat or 0.03 ha of banana or 0.06 ha of mango for satisfying once need. Thus
mango produces about 9 times more food energy than the wheat produced per unit area.
6. Raw Material for Industries: Fruit farming is the base for several industries like canning,
essential oils etc which in turn provide work for more people.
7. Use of Undulating Lands: Fruit growing can be practiced in places where the gradient is uneven
or where the land is undulating and agronomical crops cannot be cultivated. In Konkan region, mango
and cashew are cultivated on large scales on hilly and hill back area.
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Fundamentals of Horticulture
c. Environmental Appearance
Provides health & comfort by
Cleaning the air.
Preventing erosion.
Providing shade.
Providing nutrition.
SCOPE OF HORTICULTURE
Like any other things, scope of horticulture depends on incentive it has for the farmers, adaptability of
the crops, necessity and facilities for future growth through inputs availability and infrastructure for the
distribution of produce/marketing etc.
2. Adaptability:
India is bestowed with a great variety of climatic and edaphic conditions as we have climates
varying from tropical, subtropical, temperate and within these humid, semi-arid, arid, frost free
temperate etc.
Likewise we have soils from loam, alluvial, laterite, medium black, rocky shallow, heavy black,
sandy etc., and thus a large number of crops can be accommodated with very high level of
adaptability. Thus, there is lot of scope for horticultural crops.
3. Necessity:
After having achieved the self sufficiency in food, nutritional security for the people of the country
has become the point of consideration/priority.
To meet the nutritional requirement in terms of vitamins and minerals horticulture crops are to be
grown in sufficient quantities to provide a bare minimum of 85 g of fruits and 200 g of vegetables
per head per day with a population of above 120 crores.
Good land is under pressure for stable food, industry, housing, roads and infrastructure due to
population explosion and only wasteland had to be efficiently utilized where cultivation of annuals
is a gamble due to restricted root zone and their susceptibility of abiotic stress. These lands can be
best utilized to cultivate hardy horticultural crops like fruits and medicinal plants.
At present our share in international trade of horticultural commodities is less than one percent of
total trade. Moreover, these commodities (spices, coffee, tea etc.,) fetch 10-20 times more foreign
exchange per unit weight than cereals and therefore, taking advantage of globalization of trade,
nearness of big market and the size of production, our country should greatly involve in
international trade which would provide scope for growth.
4. Export value:
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Fundamentals of Horticulture
Among fresh fruits-mangoes and grapes; in vegetables- onion and potato; among flowers, roses;
among plantation - cashewnut, tea , coffee, coconut, arecanut, and spice crops like black pepper,
cardamom, ginger, turmeric, chillies, etc., constitute the bulk of the export basket.
European and gulf countries are major importer of horticultural produce
In the recent past communication and transport system have improved, investment in food industry
has increased which will support growth of horticulture through quick deliverance and avoidance
of waste.
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Fundamentals of Horticulture
II) Biennials:
These plants usually requires two years or at least two growing seasons with more or less of a dormant
season or lasting season between two completed life cycle. Seed sown in spring or summer, and vegetative
growth is completed in first year and in the following spring, flowering and fruiting takes place. Generally
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Fundamentals of Horticulture
the period of growth is 6 to 9 months e.g. Gladioli, Dahlia. No hard and fast line can be drawn between
annuals and biennials crops like turnip, carrot, cabbage and onion are classified as biennials.
III) Perennials:
Any plant that lives more than two years is a perennial e.g. Mango, Citrus. These crops are classified in to
two groups.
i) Herbaceous:
Herbaceous perennials are those with more or less soft succulent stems. In Temperate climates the tips die
off after seasonal growth but root remains alive and produce new stem and tops on favorable conditions.
In other words their tips are annual while ground parts are perennials lie many years and are classified as:
a) Trees b) Shrubs c) Vines according to their habit of growth. a) Trees:
Trees are upright in habit and stems take the form of central axis e.g. Mango, Sapota, guava, Mandarins
etc.
b) Shrubs:
Shrubs have no main trunk but a number of erect or semi erect stems are seen but do not forms the main
frame work e.g. Hibiscus, Rose, and Lantana Acalyphya etc. c) Vine:
Both woody and herbaceous have stems which are flexible and not in position to keep their branches and
leaves erect. They either spread on the ground or require some support whether alive or man made e.g.
cucurbit vines, Grape vines, Passion fruit etc.
3. Classification of Horticultural Crops Based on Whether they shed Leaves during a Portion of
Year
i) Deciduous: Fig., Guava, Apple, Ber, Sweet cherry, Pomegranate, Grape, Mulberry, Phalsa, Almond ii)
Evergreen: Aracanut, Dates, Coconut, Pineapple, Banana, Jackfruit, Avocado, Sweet orange,
Mandarin orange, K. lime, Mango, Chicku, Papaya, Passion fruit, Cashew nut
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Fundamentals of Horticulture
Tropical plants are those which do not tolerate severe cold but can tolerate warm temperatures of about
1000F. Those plants need strong sunshine, warms, humidity and a very mild winter. They cannot stand far
against frost e.g. Papaya, Banana and Pineapple.
iii) Sub - Tropical:
Sub - tropical plants like Orange, Litchi, Fig, Mango and cashewnunt are intermediate in character. They
need warmth and humidity and can tolerate mild winters.
The above classification, based on climatic preference of plants, is more or less arbitrary and no sharp line
can be drawn between these several groups. It however, indicates the broad difference in climatic needs of
various plants. This does not necessarily mean that a plant belonging to one zone does not grow in other
zones. For instances, annual crops of the temperate region like potato, knolknol and cabbage grow in
tropical and sub - tropical regions also, but they come up well only in the winter season than other climatic
zones.
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Fundamentals of Horticulture
6. Classification of Horticultural Crops based on Use of Horticultural Plants:
Vegetable:
a) Vegetables Grown for Aerial Portion:
1. Cole Crops: Cabbage, cauliflower
2. Legume Crops: Peas and Beans
3. Solanaceous Crops: Tomato Brinjal
4. Cucurbits: Cucumber, Red Pumpkin
5. Leafy Vegetables: Spinach, Methi
6. Salad Vegetables: Lettuce, Brocoli
7. Corn Vegetables: Sweet corn and Popcorn
b) Vegetable Grown for Underground Portion:
1. Root Crops: Beet. Carrot
2. Tuber Crops: Yam, potato.
3. Bulb Crops: Onion and Garlic
ii) Fruits
a) Temperate (Deciduous fruits):
1. Small Fruits: Grape, Strawberry
2. Tree Fruits: Apple, pear, Cherry
3. Nuts: Peach, Walnut
b) Tropical and Sub Tropical:
1. Herbaceous Perennials: Pineapple, Banana
2. Tree Fruits: Mango, Papaya
3. Nuts: Cashewnunt, Aracanut
II) Ornamental Plants 1.
Flowering Trees: Gulmohar, Neelmobor, Cassua
2. Road Side Trees: Neem, Baniyan tree, Rain tree
3. Shade Giving Trees: Rain tree, Mahogany
4. Flowering Shrubs: Nerium, Hibiscus, Tagar
5. Foliage Shrubs: Thuja, Casurina
6. Climbers and Creepers: Petrea, Bignonia, Ipomea
7. Bulbous Plants: Canna, Caladium, Tuberose
8. Hedge and Edges: Duranta, Clearadendron, Ageratum
9. Annuals: Pitunia, Ainnia.
10. Perennials: Chrysanthemum, Roses.
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Fundamentals of Horticulture
CLASSIFICATION OF FRUITS
Classification is the system of grouping or placing of individuals according to nomenclature. It i s very
useful to the pomologist. It helps to:
To identify and naming the crop.
To study the close relationship.
To know their hybrids and crossing behaviour.
To know their compatibility & inter grafting ability.
To know their adoptability to soil & climate.
POMOLOGY: Pomology is a branch of horticulture which deals with study of various aspects of fruits
loke, rising of saplings, growing them properly and providing various intercultural operations.
The term pomology is a combination of two Latin words means Fruits and
study .
in Greek also meaning fruits later subsequently in Latin word means fruits, logos-
study.
Basic Pomology: Study of basic aspects of fruit pro duction like training, water management, use of
In this classification, the fruits trees are categorized into three recognized groups.
i. Temperate fruits:
Temperate fruit plants are exacting in their climate requirement.
They are grown only in place where winter is distinctly cold, require as exposure of specific chilling
temperature for certain period without which they do not flower.
These fruit plants are generally deciduous and stand frost. Eg. Apple, almond, peach, pear, plum,
strawberry, apricot, persimmon, cherymoya, walnut, peanut, hassle nut, cherry, pistachios and kiwifruits
etc.
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Fundamentals of Horticulture
ii. Tropical fruits:
Tropical fruit plants are generally evergreen and are extremely sensitive to cold.
They do well under lesser fluctuations of diurnal temperature, light and dark periods they require a moist
warm climate but are capable of withstanding dry weather in some cases Eg; mango, banana, papaya,
sapota, etc.,
They are also quite adoptive to fluctuations of light and dark period during day and night.
Some subtropical fruit plants require chilling for flower bud differentiation.
B. Classification based on bearing habit: On the basis of bearing habit, fruit trees are classified in to six
categories to facilitate cultural operation like pruning, skiffing, heading back etc.
1. Fruit buds bore terminally and giving rise to inflorescence without leaves e.g. Mango, Cherry, etc.
2. Fruit buds borne terminally and unfolding to produce leafy shoots which terminate in flower clusters.
e.g. Apple
3. Fruit buds borne terminally and unfolding to produce leafy shoots with flower or flower clusters e.g
Guava
4. Fruit bud borne laterally containing flower parts only and giving rise to inflorescence without leaves or
leaves present, they are reduced in size., e.g. Citrus
5. Fruit bud borne laterally and unfolding to produce leafy shoots terminally in flower clusters this type of
flowering is noticed in grapes and cashewnut.
6. Fruit buds borne laterally and unfolding to produce leafy shoots with flower clusters in leafy axils.
eg.Fig.
C. Fruit morphology: 1) Simple fruit - Berry: Banana, Papaya, Grape, Sapota, and Avocado
2) Modified berry-
i. Balusta : Pomegranate
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Fundamentals of Horticulture
ii. Amphisarca : Woodapple, Bael
iii. Pepo : Water melon
iv. Pome : Apple, Pear, Laquat
v. Drupe (Stone) : Mango, Pear, Plum
vi. Hesperidium : Citrus
vii. Nut : Cashew, Litchi, Walnut, Rambutan
viii. Capsule : Anola, Carambola
3) Aggregate fruits : Etario of berries Custard apple, Raspberry
4) Multiple fruit : Syconus- Fig : Sorosis- Jackfruit, Pineapple, Mulberry
Mango, Banana, S apota, Guava, Papaya, Citrus, Grape, Pomegranate Pineapple Litchi,
Apple, Fig, Peach, Pear, Plum, Annona, Ber, Jamun, Cashew, Cucumber, Cherry,
Tomato Strawberry.
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Fundamentals of Horticulture
Datepalm, Ber, Amla, Guava, Pomegranate, Cashew, Fig, Mango, Apple, Citrus, Pear,
Coconut, Khirni Jamun, Phalsa Straw berry
H. Based on longevity :
a) Very Long longevity - >100 yrs- Datepalm, Coconut, Arecanut
b) Long longevity - 50-100yrs - Mango, Tamarind
c) Medium - 10-50 yrs - Litchi, Guava,Pomogranate
d) Short - Pineapple, Banana
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Fundamentals of Horticulture
Indian wild Fragaria Rosaceae India Etaerio of Fleshy thalamus & X=7
strawberry vesca achenes numerous achenes
2n=2x=14
Peach Prunus Do Do Do Do Do
persica
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Fundamentals of Horticulture
Grape fruit Citrus Rutaceae West Indies Hesperidium Juicy placental hairs X=9
paradisi
2n=2x=18
Pummelo Citrus grandis Rutaceae South east Hesperidium Juicy placental hairs X=9
asia
2n=2x=18
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Fundamentals of Horticulture
Banana Musa Musaceae South east Berry Meso & endocarp X=11
acuminata asia
2n=3x=33
4x,2x
Indian Ziziphus Do Do Do Do
jujube/ ber mauritiana
Fig Ficus carica Moraceae West asia Synconus Fleshy receptacle X=13
2n=2x=26
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Fundamentals of Horticulture
2n=2x=30
kaki 2n=6x=90
Quince Cydonia Do Do Do Do Do
oblonga
Grape Vitis vinifera Vitaceae Black sea to Berry Pericarp& Placenta X=19
Caspian sea
2n=2x=38
20
2n=58,174
Olive Olea europaea oleaceae Mediterran Drupe Epi & Mesocarp X=23
ean region
2n=2x=46
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Fundamentals of Horticulture
Climate is one of the important complex factors which influence the fruit production. It includes
several basic environmental conditions such as temperature, rain fall, humidity, and lights. Fruits
growing zones are based on the climatic factors. These zones are tropical, sub- tropical, temperate
and arid zone. However, there are certain exceptions to this, e.g. grape can be grown in temperate
and sub-tropical regions, while papaya can be grown in tropical and sub- tropical condition.
Temperate Fruits:
This class of fruits grown successfully in cold regions where temperature falls below freezing point
during winters. During the cold season, the trees, sheds their leaves and goes into rest period. For
breaking this rest period, a definite chilling period is required. This class of fruits includes fruits like
apple, pear, walnut, almond, plum, peaches, strawberry etc.
Tropical Fruits:
This class includes fruit crops which are unable to endure cool temperature. Some of them are being
severally injured by even the temperature somewhat above the freezing point. This class of fruits
required hot and humid climate in a summer and mild winter. This class includes only evergreens
such as mango, Sapota, papaya, cashew, pineapple, banana etc.
Sub-Tropical Fruits:
This class includes fruit crops intermediate in character to tropical and temperate. These fruits
grown mostly in plains where the climate is hot, comparatively dry and winter less severe. This class
of fruits includes fruits like citrus, phalsa, fig, pomegranate etc.
Arid Zones:
Besides these zones, the arid region has extreme climatic conditions. In arid zones rain is very
greatly low and its distribution is erratic leading to low plats stand and productivity. To make it
worse, the water storage capacity of soils in these areas is very low, being mainly poor textured and
shallow soils, and is compared with high evaporative losses. This class of fruits includes ber.
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Fundamentals of Horticulture
The soil is also a living system with millions of microbes that breakdown organic matter
and builds it again.
Microbes are essential and survive only when soil is well aerated and rich in organic
matter and devoid of waterlogged conditions.
Texture of soil depends on the size of solid particles and classified as gravel, coarse and
fine sand, silt and clay.
Soils are classified according to relative distribution of these particles and there are 12
textural classes.
Likewise, arrangement of these particles is referred as structure, and both texture and
structure lend soil physical properties like water holding capacity, aeration and bulk
density.
Generally loamy soils and crumb structure are most pre ferred for fruit crops.
According to level of organic matter, soils are classified as mineral soil or organic soil
and soil having more than 20% organic matter is organic soil like peat and muck.
Minerals and salts lend chemical properties to the soil like pH, alkalinity, sodicity,
salinity and cation exchange capacity which influence the availability of nutrients in soil.
Therefore, for making choice for soil, soil analysis in terms of following criteria is
essential to decide on land capability.
Criteria for land capability class:
i. Slope and erosion hazard.
ii. Soil depth.
iii. Drainage. iv. Workability.
v. Stoniness and rockiness.
vi. Water holding capacity.
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Fundamentals of Horticulture
vii. Permeability.
viii. Nutrient availability.
ix. Fertility status.
Drought tolerant crops: Ber, Aonla, Ahalsa, Lasoda, Kair, Custard apple, Karonda, Fig, Guava
etc.
Scrapping of salts and reclamation of soil by application of gypsum, iron pyrites, press mud etc., on regular
basis in case of salinity problem.
Replenishment of nutrients harvested by the crop on regular basis by preparing a balance
sheet for nutrients.
Soil is the most important natural resource for fruit culture and it needs to be protected and improved.
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Fundamentals of Horticulture
CLIMATE
Climate is the most important factor on which choice of the crop for a region depends and
therefore, understandi ng about soail and climate and their requirement for different crops
for optimum production on sustainable basis is important for horticulturists.
Climate is defined as the whole of average atmospheric phenomena for a certain region
calculated for a period of thirty years. These phenomena are light, heat, water and air. LIGHT:
Electromagnetic radiation to which the organs of plant react ranging in wavelength from
4000 to 7700 angstrom units, and is propagated at a speed of about 540 kilometres per
second.
It is essential for the process of photosynthesis and therefore, for growth and
development of plants.
There are two aspects of light, its intensity and duration which are important for plant
development.
The light intensity can be estimated from the number of hours of bright sunlight or from
the cloudiness of sky.
Generally horticultural crops need a lot of light and must be grown in sunny climate, but
there are some crops which can tolerate shade. Eg. Turmeric and ginger.
There are others like youn g mangosteen, coffee, cocoa and tea need shade during part of
their development.
A third group requires permanent shade like salak palm, duku, and carambola.
The duration of light for the time elapsing between dawn and dusk referred as photoperiod
or day length. This exerts considerable influence on flowering.
Based on the response by plants the major classes are following. However, fruit crops for such
categories are not known.
1. Long day plants: Cabbage, Cauliflower, Onion, Beetroot, Radish, Carrot, Spinach, Potato, Dill and
Plantago.
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Fundamentals of Horticulture
2. Short day plants: Strawberry, Pineapple, Chrysanthemum, Poinsettia, Aster, Balsam, Salvia,
Euphorbia and Xanthium.
3. Day neutral plants: Tomato, most fruit cro ps, Pepper, Cucumber, Snapdragon, Mirabilis and
certain varieties of peas.
HEAT:
Heat is a non-mechanical energy transfer with reference to a temperature difference
between a system and its environmental surrounding.
Availability of heat units decide the crop for a given place and the average temperature of
a place gives an idea about heat units available on the basis of which crop can be decided.
Temperate fruit crops like apple, pear, peach, plum and almond become dormant due to
short day conditions in the region and need chilling of various lengths to break dormancy.
Frost and chilling are harmful for tropical and subtropical plants.
On the other hand extremely high temperatures found in arid region cause wilting, sunscald,
necrotic spot and even death of plants.
Therefore, under such conditions appropriate choice of plants and provision of protection
become important. Based on temperature v ariations on the surface of the earth we have the
following climates.
Tropical equable climate with no distinct winter.
Subtropical Climate with distinct winter and summer.
Temperate: Distinct winter, summer and autumn with temperature below freezing during winter is
common.
1. Tropical : Mango, Banana, Papaya, Sapota, Pineapple, Coconut, Cashew, Arecanut, Breadfruit,
Jackfruit and Avocado.
2. Subtropical: Guava, Grape, Citrus, Date palm, Phalsa, Pomegranate, Litchi and Loquat.
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Fundamentals of Horticulture
3. Temperate: Apple, Pear, Peach, Plum, Quince, Apricot, Walnut, Almond, Strawberry and
Cherry.
WATER
Water is a transparent, odourless and tasteless liquid compound of hydrogen and oxygen
(H20) with 11.91% hydrogen and 88.81% oxygen.
Near equator the total rainfall is 2000 mm per year and away from it, which reduces but
again influenced by a number of factors like mountain ranges.
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Fundamentals of Horticulture
Whereas high humidity favours fungal diseases. Plants liking for high humidity and low
humidity are there:
Low humidity (Dry) : Ber, Grape, Date palm, Pomegranate, Citrus, Aonla and Guava.
AIR
A mixture of oxygen, nitrogen and other gases that surrounds the earth and forms its
atmosphere.
If its quality is polluted by the accumulation of gasses like hydrocarbons, SO2, CO2, CO,
NO, ethylene and methane the plant growth adversely affected but we are more
concerned with the movement of air (wind) causing great damage to crops in deserts,
coastal areas, valleys for w hich provision of windbreaks and shelterbelts are suggested
and such situations sometimes have to be avoided for plantation.
Storm has a wind speed of 50/hr whereas, hurricane has a wi nd speed of more than
100km/hr.
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Fundamentals of Horticulture
Objective of propagation: To produce individuals those are identical to the mother plant or original
plant. Thus a successful propagation method is one which transmits all the desirable characteristics
of a mother plant to the offspring.
In general field crops as well as almost all the vegetables and flower crops are commercially
propagated by seeds but fruit crops are mainly propagated by vegetative means of propagation.
Normally, field crops, most of the vegetables and flower crops are highly self pollinated in natures
which are considered as homozygous. Hence when they undergo sexual reproduction, the resultant
off springs will be homozygous in nature, similar to the mother plant. But in case of fruit crops, they
are highly cross pollinated in nature, means highly heterozygous. Hence on sexual process they will
produce off spring- not true to the type of mother plant. Therefore, in fruit crops in general vegetative
means of propagation is more desirable to get true to the type of mother plant than sexual method
of propagation.
Sexual propagation: Raisings of plant by means of seed. It has certain advantages over vegetative
means of propagation-
Seedis relatively a cheaper and simple way of obtaining large number of plants as compared
to vegetative propagation.
Seedlings have usually long life as compared to vegetatively propagated plants.
Seedling plants have better root system and therefore provide better anchorage than
vegetatively propagated plants.
It is the only practical method of propagation of most of the vegetables, annual flowers and
fruit plants like papaya, phalsa, mangosteen which cannot be propagated by vegetative
means.
Hybrids are 1st raised through seeds.
Rootstocks for budding and grafting purpose are raised through seeds only.
Sexual propagation may sometimes leads to the production of chance seedlings which may
be superior to the mother plant.
Seedling plants are comparatively more resistant to insect-pest and diseases than the
vegetatively propagated plants.
Seeds can be stored for a longer time and can be easily transported to distance market.
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Fundamentals of Horticulture
Seedlings are usually hardy and can tolerate adverse climatic conditions in a better way than
the vegetatively propagated plants.
No special technical skill is required for raising plants through seeds.
Majority of viruses are not transferred through seeds. Thus, seed propagation is useful in
producing virus free plants.
Nucellar seedlings can be utilized to raise uniform plants.
In polyembryonic crops like mango, citrus seeds on sowing give rise to more than one
seedling. Thus seed is the desirable means of propagation in such crop.
Mangosteen fruit is developed through parthenogenesis. Seedlings obtained from seeds of
such fruits are similar to its mother plant and thus propagated commercially through seed.
Disadvantages:
Seedling plants are highly heterozygous in nature. Owing to segregation, the seedling trees
are not uniform in growth, yield and fruit quality parameters.
Seedling plants are usually tall and spreading types, thus the management cost involved in
operation like harvesting, pruning and plant protection etc. is comparatively higher.
Seedlings plants have usually long juvenile phase and take more time to come in bearing.
In case of recalcitrant crop, seed loose viability very soon after extraction and thus very low
germination rate.
The beneficial effect of rootstock on scion variety cannot be taken in sexual propagation.
Some fruit plants like pineapple, banana does not produce viable seeds and propagation
through seeds is not possible in them.
It is not possible to perpetuate the exact characters of any superior selection through seeds.
Classification of seed on the basis of storability: On the basis of storability of seeds in relation to
moisture content, they are classified into two groupsviz. Orthodox seeds and recalcitrant seeds.
a. Orthodox seeds: The seedswhich can be dried to low moisture level(5-8% of lower) and
lose their viability with the increase of moisture, are called as orthodox seeds. The viability
of these seeds can be maintained by drying the seeds and storing them at low temperature
for longer period of time. Eg. Ber, custard apple, date palm, fig, grape, guava, mulberry,
papaya, passion fruit, peach, pineapple, plum, phalsa, pomegranate etc.
b. Recalcitrant seeds: The seeds, which remain viable at relatively higher moisture level (8-
15% or above) and if dried below certain moisture level, they start to lose their viability, are
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Fundamentals of Horticulture
called as recalcitrant seeds. Eg. Avocado, Barbados cherry, carambola, bread fruit, jack fruit,
litchi, mango, mangosteen, rambutan, citrus etc.
Vegetative propagation: Any plants are propagated commercially through asexual mean in which
vegetative parts of the plant are used. It is possible because all the living cells of a plant have a
capacity to regenerate into a full plant under favourable environmental condition. This tendency of
plant cell is called as totipotency. The term given by Haberlandt in 1902. The plants produced by
vegetative means are therefore genetically identical and similar to the mother plant.
Advantages:
True to the type and uniform in growth, yield and fruit quality
Ideal method of propagation for the plants which are more prone to seed dormancy (all the
temperate fruits).
Vegetatively propagated plants are less vigorous and thus can easily be maintained as
compared to seedling plants.
Some fruit crops like banana, pineapple, fig and varieties of grape, guava and lemon produce
seedless fruit and have no viable seeds. Thus can only be perpetuated through vegetative
means.
Plants come in bearing earlier than seedling plants.
Possible to regulate the tree size, fruit quality and precocity in bearing by exploiting the
desirable effects of different rootstocks.
It is possible to exploit the desirable abiotic effect of rootstocks on scion cultivar by budding
and grafting.
Helpful to overcome the problem of self incompatibility of different fruit crops by top
working the desirable pollenizers on scattered trees throughout the orchards.
Detection of virus in the plant system by using indicator plant is only possible through
budding or grafting. Eg. Kagzi lime can be used as rootstock (indicator plant) for detecting
the presence of tristeza virus in citrus.
Helps in shortening in breeding cycle by grafting scion of new cultivar on to a large
established tree or on certain dwarfing rootstock for early assessment.
A clone can only perpetuated by vegetative means.
Benefit of certain interstock can only be obtained through vegetative means.
Damaged part of the tree trunk or root can only be repaired by bridge grafting or inarching.
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Fundamentals of Horticulture
Disadvantages:
Parthenium, raspberry, apple, onion, Poa etc. In some species, this phenomenon occurs without the
stimulus of pollination whereas in others, pollination appears to be necessary for the development
of a viable embryo.
b. Non-recurrent apomixis: Embryo develops directly from the haploid egg cell or some other
haploid cells of embryo sac without fertilization and as a result, the embryo developed is also
haploid in nature. It is common in Solanum nigrum and Lilium species.
c. Adventitious embryony: It is also called as nucellar embryony. Here the embryo does not
develop from the cells of the embryo sac but from a cell or a group of cells either of nucellus
or integuments. Such embryos usually develop outside the embryo sac in addition to the
regular embryo. This is quite common in citrus, where normal pollination and fertilization
takes place in usual manner and apomictic embryos are develop outside the embryo sac.
d. Vegetative apomixis: In this type, vegetative buds or bulbils are produced in the
inflorescence in place of flowers. These buds or bulbilis may sprouts into new plants while
they are still attached to the mother plants. This is quite common in Allium, Agave, Poa and
Dioscorea.
Apart from these, different scientists also included polyembryony as one type of apomixis. But it is
not a separate type of apomixis. It is the phenomenon in which two or more embryos are produced
in a single seed. The condition may results from many reasons. One of the most common reasons
being the nucleus develops within the embryo sac, which may lead to the development of more than
one embryo. Cleavage of pro-embryo during the early stages of development may the other reason
for the development of multiple embryos.
Significane of apomixis:
Produce homozygous line
Useful for producing uniform rootstock. Eg. Rootstock of apple like M. tiringoides, M.
hupehensis, M. sikkimensisetc.
Apomictic seedling are quite healthy and highly uniform
Effective for the production of virus free quality planting materials
Cuttings
Types of cuttings
A. Stem cutting: Next to seed, stem cuttings are the most convenient and popular method of plant
propagation. Most the cutting techniques fall into this category. A stem cutting is any cutting taken
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Fundamentals of Horticulture
from the main shoot of a plant or any side shoot growing from the same plant or stem. There are few
general considerations which helps in selection of suitable cuttings. First of all it is essential for the
cutting to have sufficient reserve food to keep tissues alive until root and shoots are produced. The
shoots with high carbohydrate content usually root better. To maintain high carbohydrate content in
a shoot, ringing or notching stem down to the wood are useful practices. As a general rule, cutting
from young plants root better but if older shoots of the plants are cut back hard, very often they can
be induced to produce suitable shoots for rooting. Broadly there are four types of stem cuttings
which are as follows-
i. Hardwood cuttings: It is simple method of plant propagation in which cuttings are made from
the mature and lignified stems of shrubs and trees. Such cuttings are easy to secure and can be
easily handled, stored and transplanted. This type of cutting is prepared during dormant season.
Usually from one year old immature shoots
selected and weak, fast growing shoots with long internodes should be avoided. The length of
the cutting varies from 10-45 cm in length and 0.5-2.5 cm in diameter, depending on the species.
usually the cutting of 25-30 cm length with pencil thickness are preferred. Each cutting should
have at least 2 buds. While preparing the cutting, a straight cut is given at the base of shoot below
the node while a slanting cut, 1-2 cm above the bud is given at the top of the cutting. This helps
in maintaining the polarity of the shoot and if rain occurs, water does not accumulate on the tip
of the cutting, which saves the cutting from fungal infection. It also helps in maintaining polarity
of the cut sticks. It is often advantageous to take hardwood cutting with a heel (in temperate fruit
crops), that is with a piece of old wood, attached to the base. Presence of Mallet (present in
Quince) at the base of cutting should also be preferred. Similarly, stem cutting will more often
root better if bases are etiolated, notched or girdled or ringed before being removed from the
parent plant. After preparation of cutting, the basal portion of those cutting should be treated with
root initiating hormone, especially Indole-3-butyric acid (IBA). For deciduous crop, the dose of
IBA varies from 2500-5000 ppm (with maximum 10,000 ppm in difficult to root crops) while in
evergreen plants it is 2000 ppm to slightly higher (5000-10000 ppm in difficult to root crops). It
is generally practised in a number of deciduous fruit plants like grape, hazelnut, chestnut,
fig, quince, pomegranate, mulberry, plum, olive are commonly propagated by hardwood cutting.
ii. Semi-hardwood cutting: The semi-hardwood cuttings are prepared from partially matured,
slightly woody shoots. These are succulent and tender in nature and are usually prepared from
growing wood of current season growth. Usually those shoot, which snap clean when broken, are
considered ideal for preparation of semi-hardwood cutting. The length of the cutting varies from
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Fundamentals of Horticulture
7- 15 cm. The cuttings are prepared by trimming the branches with a straight cut below the node
and removing a few lower leaves at the base and retaining 2-4 leaves at the top of the cutting.
Treating the cutting with IBA @ 1000- 3000ppm (maximum 5000 ppm in difficult to root crops)
before planting is beneficial for commencement of proper rooting. While planting one quarter of
their length should be inserted in the soil. The best time for taking cutting is late spring to summer,
when new shoots have emerged and their wood is partially matured.
It is normally practised in evergreen fruit plans like mango, guava, lemon, jackfruit, olive etc. iii.
Softwood cutting: Softwood cutting is the name given to any cutting prepared from soft, succulent and
non-lignified shoots which have not become hard and woody. Usually, the cutting size is 7.5- 12.5 cm
but it varies from species to species. Usually some leaves should be retained with this type of cutting.
Before planting treatment with IBA (500-1250 ppm with maximum of 3000 ppm in difficult to root
crops)is beneficial. The best time for the preparation of softwood cutting is spring to early summer.
It is normally practiced in different flowering annuals like Juniper, hollyhick, lilac. But some time
it is also practised in apple, pear, peach, plum etc.
iv. Herbaceous cutting: The terminal leafy portion of the stem of a plant is used for preparing
herbaceous stem cutting. These cuttings are soft and succulent with length varying from 7.512.5 cm.
These cuttings are rooted under the same conditions as the softwood cuttings, requiring high relative
humidity but are liable to wilt soon if proper humidity is not maintained. Therefore, much attention
is required for working with herbaceous cuttings. Herbaceous cuttings of some plants exude a sticky
sap that interferes with rooting process. In such cases basal end of the cutting should be allowed to
dry for few hours before planting. For proper root initiation treatment of the basal portion of the
cutting with IBA @ 500-1250. It is generally practiced in flowering annuals like Geranium,
poinsettia, dieffenbachia, chrysanthemum. In fruit crops it is followed in pineapple (slips and
crown), and in persimmon.
B. Root cuttings: Propagation by means of root cuttings is also a simple and cheap methods of
vegetative propagation in species which are difficult to propagate with other methods. In general the
plants which produce suckers freely are easily propagated by root cuttings. For preparation of root
cuttings, roots which are of 1 cm thick and 10-15 cm long are cut into pieces. The best time for
taking root cuttings is late winter or early spring when the roots are well supplied with stored food
materials but before the new growth starts. However in temperate fruits, root cuttings are prepared
in the month of December and are kept in warm place in moss grass or wet sand for callusing and
are then transplanted during February- march in open beds. While using root cuttings, it is important
to maintain the correct polarity. Thus, to avoid their planting upside down, the proximal end of the
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Fundamentals of Horticulture
cutting should have straight cut and distal end a slanting cut. The proximal end should always be
kept up. While planting, insert the cuttings vertically so that the top is above the soil level. However,
in some cases, horizontal planting gives satisfactory results as in sweet potato.
Balckberry and raspberry are commercially propagated by this method. However, kiwi fruit, bread
fruit, fig, mulberry, apple, pear, peach, cherry and persimmon are also propagated by root cuttings.
Apart from theses different horticultural crops particularly different flowering annuals and vegetable
crops also propagated by leaf cutting due to their herbaceous nature but fruit crops are rarely
propagated by this particular means of propagation. However, leaf bud cutting technique is
sometimes followed for the propagation of black berry, raspberry, lemon.
Budding
In grafting, the scion is a detached piece of shoot or stem with several buds. But in budding, scion
consists of only one bud and small portion of wood. Thus, budding is a form of grafting and often
called as bud grafting.
Advantages:
Quick and efficient method of propagation as compared to grafting.
Best propagation method if propagating material is scarce.
Budding is useful in plants release excessive wound gum (stone fruit) from injury carried to
xylem or wood portion of stem at the time of grafting.
Budding results into stronger union compared to grafting thus injury to the budded plants due to
storms and strong wind is less.
It is comparatively simple method of plant propagation than grafting and can be done by common
peoples.
Types of budding
i. Shield or T budding: as the name indicates shield is the shape of the bud and T is the shape
of cut given on the rootstock for the budding operation. It is the most common method of
budding practiced by nurserymen throughout the world. For T budding, one year old rootstock
seedling of 25-35 cm height and 2-2.5 cm thickness is selected which is in actively growing
stage, so that the bark will separate readily from the wood. In this budding, 1 st a vertical cut
(2.5 cm length) to be made on the stock at 0.-2.5 cm above the soil level. Then the horizontal
crosscut to be made at the top of the vertical cut to give T shape. As the horizontal cut is made,
the knife is given a twist to open the flaps of bark for the insertion of the bud. Thereafter, scion
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Fundamentals of Horticulture
bud is prepared by removing the bark shield with a bud. For this, an upward slicing cut is
started at the point on the stem at 12-13 mm below the bud, continuing under the bud to an
about 2.5 cm above. The shield piece should be thin, thick enough to have some rigidity. A 2nd
cut is then made at the upper portion of the 1 st cut which permitting the removal of the shield
piece. After removal of the scion bud, it should be inserted into the T cut on the stock by
pushing the shield downward under the two raised flaps of bark until its upper horizontal cut
is matches the same cut on the stock. Then wrapping of the bud union must be done using
budding tape.Eg. Apple, pear, peach, plum, apricot, cherry, rose and citrus are propagated by
this method.
ii. Inverted T budding: In area that experiencing high rainfall during budding season, water
running down the stem of the rootstock may enter the T-cut, soak under the bark of the stock
and prevent into the shield piece from healing into place. In such condition, inverted T budding
gives better results, since excess water is shed. For this, rootstock has the transverse /horizontal
cut at the bottom rather than at the top of vertical cut. In removing the shield piece from the
bud stick, the cut on bud stick should start above the bud and continuing downward below it.
Then insertion of shield into rootstock should be done from lower part to upward direction.It
is usually practiced in citrus.
iii. Patch budding: Widely successful in thick barked species such as walnut, pecan nut. In patch
budding, bark of both stock and scion requires to slip easily. A special knife called double
bladed knife is used in this budding technique which makes two transverse parallel
cut 2.5-3.5 cm apart through the bark to the wood in a smooth area of the rootstock about 10
cm above the ground. Then two transverse cuts are connected at each side by vertical cuts made
with a single bladed knife. The patch of bark containing a bud is cut from the budstick in the
same manner as of rootstock. The size of the bark removed from the budstick should be same
as of size of the bark removed from the rootstock. After the removal of bud patch from the
budstick, it must be inserted immediately on the already prepared rootstock. The inserted patch
is now ready to be wrapped.
iv. I- Budding: In this budding, two parallel cuts are made on the rootstock using the same parallel
double bladed knife, used in patch budding. These two transverse cuts are then joined at their
c
then be raised to insert the bud patch beneath them. A better fit may occurs if the side edges of
the bud patch are slanted. It is mainly practiced in ber.
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Fundamentals of Horticulture
v. Ring budding: In ring budding, a complete ring of bark is removed from the stick and it is
completely girdled. A similar ring of bark containing a bud is removed from the bud stick and
it is inserted on to the rootstock. In ring budding, both stock and scion should be of same size.
However, it is applicable in stocks having thickness below 2 cm. It has been utilized in ber,
peach, mulberry because the newly emerged shoots from the heavily pruned plants are capable
of giving such buds for budding, which can easily be separated. In this method, since the stock
is completely girdled and if the bud fails to heal in, the stock above the ring may eventually
die.
vi. Flute budding: In flute budding, the bark patch from the stock is removed in such a way that
it must completely encircles the stock except with a narrow bark connection between the upper
and lower cuts on the stock. A similar patch of bark is removed from budstick containing a
healthy bud. The shield containing the bark is then inserted in the vacant area in the stock.
After fitting it properly and tightly on the stock, it should be wrapped with suitable wrapping
material, leaving the bud uncovered. Because of the presence of narrow connecting strip of
bark on the stock, it remains alive even if the bud fails to sprout. It is mainly practiced in ber,
mulberry, peach etc.
vii. Chip budding: it is a successful method of budding when the bark of the rootstock does not
slip easily and plants are not in actively growing condition.In this technique, a chip of bark is
removed from a smooth place between nodes near the base of the rootstock and replaced by
another chip of the same size and shape from the budstick, which contains a bud of the desired
cultivars. The chip in both budstick and rootstock are cut out in the same manner. In the
budstick, the 1st cut is made just below the bud and down into the wood at an angle of 30-45
degree. The 2nd cut is started about 25 mm above the bud and goes inward and downward
behind the bud until it intersects the 1st cut. The chip is removed from the stock in
the same manner and replaced by the one from budstick. After proper fitting of bud piece on
the stock wrapping should be done properly with suitable wrapping material, leaving the bud
uncovered.Mainly followed in grape, apple and pear.
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Fundamentals of Horticulture
Lecture 7: GRAFTING
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Fundamentals of Horticulture
then inserted into the split portion of the stock. A common mistake in cutting scions for this type of
grafting is to make the cut on the scion too short and the slope too abrupt, so that the point of contact
is only at the top. Slightly shaving of the sides of the split in the stock will permit a smoother contact.
After preparation of the scion, they are inserted into the stock properly and wrapped the stock tightly
with poly grafting tape or adhesive tape at the point of graft union to hold the scions in place more
securely. Thereafter, thorough waxing of the completed graft is essential.
Mostly used for rejuvenating the old orchards of the temperate fruits by top working. Apart from
this, Walnut, hazelnut, pecan nut, grape are also propagated by this method.
d. Wedge grafting: A heavy, short bladed knife is used for making a V-wedge in the side of
the stub, about 5 cm long. For this two cuts are made, coming together at the bottom and as
far apart at the top according to the diameter of the scion. These cuts extend about 2 cm deep
into the side of the stub towards the centre. The diameter of the stock to be grafted is the
same as cleft grafting- 5-10 cm and the scions are also the same size- 10-13 cm long and 10-
13 mm thick. After preparation of the stock, the scions are prepared by trimming the base to
a wedge shape, exactly the same size and shape of the V- wedge on the stock..in a stock of
5 cm diameter, 2 scion should be inserted while in stock of 10 cm diameter, 3 scions should
be inserted. After all the scions are firmly inserted into the place, all cut surfaces including
the tip of the scion should be waxed thoroughly. Like cleft grafting, it can be made in late
winter or in early spring before the bark begins to slip.
Mainly use to topwork the old orchards.
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Fundamentals of Horticulture
It refers to the production of plants from very small plant parts, tissue or cells under aseptic (in vitro)
condition. It is mainly based on the principal of totipotency. Strawberry was 1st commercially
propagatedby micropropagation technique
Advantages:
1. large scale multiplication of virus free panting materials in limited time & space
2. Through out the year production
3. Beneficial in plants where vegetative propagation is not easy like papaya
4. Production of secondary metabolites
5. Long term storage in lesser space
6. Production of seedless variety through embryo rescue (Grape)
7. Shortening of breeding cycle
8. Homozygous plant production
Methods of micropropagations
Meristem tip culture: Itconsist of 1 or 2 pairs of leaf primordia. Helpful for the
elimination of virus from infected plant.
Callus culture: Unorganised mass of parenchymatous cells. Organogenesis in two steps -
st nd
1 formation of meristem and 2 active growth of stem bud and root
Anther culture: produce homozygous plants. 1sy t time reporte d by Guha &
Maheshawari (1959) in Datura. Stage - near microscopic mitosis, when pollen grain pass
through uninucleate to binucleate condition.
Cell suspension culture: Homogenizing a piece of callus into liquid medium and
shaking with shaker until medium b ecome cloudy with suspended cells .
Ovule culture: Unfertilized or just fertilzed ovule excised and cultured. Helpful to
obtained plant in self compatible species
Embryo culture: Embryo excised at immature stage but before their degeneration
Different stages of micropropagation
Shoot initiation
Shoot multiplication
Shoot elongation
Rooting
Acclimatization
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Fundamentals of Horticulture
Applications
In India, the technique has been perfected in banana, grape and papaya
Banana: Shoot tip excised from rhizomes of sword suckers
Grape: Shoot tip with two nodal micro cutting, embryo rescue
Papaya: Shoot tip
Limitations:
Costly
Require technical skill
Chances of contamination
Plants having high level of phenols (mango, datepalm, coconut) not responds properly
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Fundamentals of Horticulture
1. Polyhouse: When the green houses are covered with plastic then it is called polyhouse.
Polyhouse
2. Net house: When the glazing material in greenhouse is net then it is called net house.
Net house may be half shade net house or complete net house.
Net house
3. Plastic tunnel: It is tunnel like structure covered with plastic and smaller than playhouse. Its
cost of construction is very lesser than Plastic tunnels; therefore small income group farmers
easily may construct it. It is constructed on arch shaped steel or
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Fundamentals of Horticulture
bamboo body. Small plants like vegetables, flowers can be cultivated inside it. Sometimes
portable plastic tunnels are constructed. These portable tunnels can be kept at safer place
when they are not in use.
Plastic tunnel
4. Mist chamber: It is nursery or plant propagating structure in which water is sprayed in
the form of mist to maintain high humidity. The objective of misting is to retain a
continuous film of water on the leaves to maintain turgidity by reducing transpirational
loss. Mist chamber also prevents from disease infection in cuttings by washing off fungus
spores before they attack the tissues.Generally fine sized nozzle is used to produce very
fine mist. The water should have good pressure and it must be free from salts. The
optimum pH of water to be used in mist unit is 5.5 to 6.5. Well -drained rooting medium
is used for adequate removal of excess water.
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Fundamentals of Horticulture
Mist chamber
Plant growing structures / containers:
In nursery, seedlings or saplings are grown in containers. The containers are used for
1. Raising of seedlings
2. Raising rootstocks
3. Hybrid seed production
4.Growing potted plants
Advantages of containers in greenhouse production
1. Uniform production of planting materials
2. Quality production
3. Easy to maintain
4. Reduce production time
C. Store house: It is very necessary for storing of fertilizers, impl ements, tools, nursery
stocks like seeds, bulbs, corms, tubers and other planting materials. A store house has
generally two compartments, one is for storing of fertilizers, implements, tools and the
other is for seeds, bulbs, corms, tubers and other planting materials which is well
ventilated. Store house is made up of brick and mortar where seed compartment contain
wooden or split bamboo shelves in several layers for storing bulbs, corms, tubers and
other planting materials
Seed dormancy refers in seeds to failure of a viable seed to germinate even when given favourable
environmental conditions. In many species can get a clue about requirements for breaking
dormancy by studying the natural environment of the plant.
Significance of Dormancy
1. From an ecological perspective, dormancy is an important survival mechanism that favours
propagation and dissemination of seeds to establish plant populations.
44
Fundamentals of Horticulture
2. Dormancy allows the seeds to remain in suspended animation without any harm during drought,
cold or high summer temperature.
3. The dormant seeds can remain alive in the soil for several years. They provide a continuous
source of new plants even when all the mature plants of the area have died down due to landslides,
earth quake, floods, epidemics or continued drought.
4. It helps the seed to get dispersed over long distances through unfavourable environment or
inhospitable area.
5. The small seeds with impermeable seed coat belonging to edible fruits come out of the alimentary
canals of birds and other animals uninjured e.g., Guava.
6. Dormancy induced by the inhibitors present in the seed coats is highly useful to desert plants.
The seeds germinate only after a good rainfall which dissolves away the inhibitors. The rainfall
ensures the seed a proper supply of water during its germination.
7. It follows storage of seeds for later use by animals and man.
Dormancy Classification
Basically it is of two types-
A) Primary dormancy: It induced during seed development resulting in seeds that are dormant
when dispersed from the mother plant.
B) Secondary dormancy: It induced by unfavourable environmental conditions following
shedding of mature seeds from the parent plant. Secondary dormancy can be induced by
conditions unfavourable for germination, such as adverse temperatures, illumination or oxygen.
Types of secondary dormancy include:
i) Thermodormancy: It is caused by prolonged exposure of imbibed seeds to temperatures
unfavourable for germination, generally high temperatures.
ii) Skotodormancy: It is induced in seeds that require light for germination when they are imbibed
in the dark for extended periods of time.
iii)Photodormancy: It occurs in seeds that are inhibited by light when they are exposed to an excess
of light.
Note, that secondary dormancy is not only induced in non-dormant seeds, but also in seeds that
already have some form of primary dormancy. For example, a lettuce seed that requires light to
germinate is said to be photosensitive and, when imbibed in the dark, it will not germinate until light
is provided. However, if dark imbibition is extended for a period of time, skotodormancy can be
induced and the seed will not germinate even if it is placed under optimum light conditions.
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Fundamentals of Horticulture
However, primary dormancy can again be classified into the following ways-
A. Seed coat dormancy
1. Physical dormancy Here the seed coats are impermeable to water with low moisture
level. Legumes are an example. Embryo is generally quiescent. The cause of physical dormancy is
the structure of the outer cell layer which becomes impermeable to water. Macrosclereid cells, a
mucilaginous outer cell layer, or a hardened endocarp are three reasons that seed coats become
impermeable to water. Such seed coats develop during the last stages of seed development.
2. Mechanical dormancy - Seed coats are too hard to allow the embryo to expand during
germination. Causes include: Structure of seed coats or remaining fruit. Occurs in olive.
3. Chemical dormancy - Presence of chemical inhibitors in the outer coverings of many fruits
and seeds. This occurs in fleshy fruits, hulls, and capsules of many dry fruits. Examples are apples,
citrus, grapes, desert plants. Very often this kind of dormancy disappears with dry storage. This
kind of dormancy may also be present in other tissues surrounding the embryo such as the
endosperm.
B. Morphological dormancy- Embryo is not fully developed at time of ripening. Need
additional embryo growth after the seed is separated from the plant. This type of dormancy mainly
observed in several herbaceous flower, seed such as Ranunculus, poppy; woody species such as
holly; tropical plants such as date palms.
C. Endogenous dormancy: Here the dormancy is controlled internally within the living
tissues of the seed. It may be exerted by semi-permiability of the seed cpvering and secondly due to
dormancy present within the embryo. It may be of two types-
i. Physiological dormancy: Most of the temperate fruits (apple, pear, peach, plum, cherry),
vegetables and flowering annuals have physiological seed dormancy lasting from 1-6 months, which
disappear on storage. The seeds of these species have physiologically immature embryo and they do
not germinate even under favourable conditions.
ii. Embryo dormancy: Here the dormancy is due to dormancy of the embryo which over
winter in soil and germinate in spring under natural condition.
D. Epicotyl dormancy: Seeds of some species have different stratification requirement for
their radicles, hypocotyls and epicotyls and this type of dormancy is called as epicotyls dormancy.
On the basis of chilling requirement for epicotyls, different plant species are divided into two sub
groups. In one group, seed germinate initially during warm period of 1-3 months and produce roots
and hypocotyls but these require 1-3 months chilling for epicotyls to germinate as in lity and peony.
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Fundamentals of Horticulture
E. Double dormancy: In certain plant species, seeds have dormancy due to hard seed coats
and dormant embryos. This is called as double dormancy. For proper germination, both the blocking
conditions must be modified to allow water to penetrate the embryo and after ripening of the
embryos. This type of dormancy is found in trees and shrubs producing seeds with hard seed coats,
grown primarily in cold winter area. To overcome the problem of double dormancy, the seeds are
first treated in a way to soften the seed coat and then cold stratification treatment is given to them.
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Fundamentals of Horticulture
- Chemical scarification involves the use of a chemical compound to degrade the seed coat. The
most common compound used for chemical scarification is sulfuric acid.
Other alternatives are sodium hypochlorite and hydrogen peroxide.
It must be noted that, despite dormancy breaking and enhancement of germination, scarification
treatments create damage to the seed due to the disruption of essential cells, favoring fungal invasion
and mechanical injury. Precautions should be taken to minimize damage while maximizing
dormancy relief. (108) For instance, optimal times for chemical scarification with sulfuric acid
should be determined for each species. Longer periods will damage the seed, while shorter periods
will be ineffective in dormancy breaking. In this example, 90 minutes in sulfuric acid were effective
to overcome physical seed dormancy in Acacia caven. Note the improvement in seed germination
of treated- versus non-treated seeds. (109) In this picture is the difference in volume of treated versus
untreated seeds, showing the lack of water uptake in non-treated seeds. While 90 minutes of
scarification in sulfuric acid are required to overcome Acacia caven seed dormancy, in the case of
Prosopis chilensis only 10 minutes was required. In this species more time would result in severe
seed damage, while 10 minutes in Acacia caven would be ineffective in dormancy breaking.
Afterripening is defined as the progressive loss of dormancy in mature dry seeds. Afterripening
rates may increase in response to environmental factors such as increased oxygen and temperatures
and decrease with increasing seed water contents. (112) In this figure, the effects of afterripening in
reducing seed dormancy of Phleum arenarium are presented. After 6 and 13 months of dry
afterripening, the seeds achieved higher germination percentages under a wider range of
temperatures.
Although the mechanisms explaining afterripening remain unknown, some possiblilities exist. For
example, in seeds with morphological dormancy, afterripening may explain the progressive growth
of the embryo during dry storage.
Afterripening in seeds with physiological dormancy could be associated with a reduction in the
concentration of a inhibitory compound or a lower sensitivity to it. For example, this table shows
how the seed ABA content from two tomato genotypes decreased after one year of dry storage.
Stratification is the exposure of the imbibed seed to low or warm temperatures. Because the most
common approach is to expose seeds to cold temperatures, stratification is often used as a synonym
for chilling or prechilling treatments. Use of warm stratification has been usually associated with
release of morphological dormancy, while cold stratification has proven effective in overcoming
physiological dormancy.
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Fundamentals of Horticulture
The temperature used for cold stratification usually ranges from 3 to 10°C, and the times of
treatments vary depending on the species. While cold stratification is an absolute requirement for
germination for some species; in others, it may only improve germination rates or uniformity. For
some species, cold stratification may increase the range of temperatures under which germination
occurs, or increase light sensitivity in species that require light for germination. (115) Effects of cold
stratification in seeds with physiological dormancy are probably the result of changes in the balance
of endogenous promoters and inhibitors. For instance, this figure shows the changes in ABA and
GA concentrations in plum seeds during stratification. A drastic reduction in the ABA concentration
and increase in GA concentration occurred that would explain dormancy breaking in this species
after stratification.
In this example, germination rates of liquidambar seeds were improved by a stratification treatment
of 15 days at 0°C, and this effect was similar to seed imbibition in a GA solution. Note that final
germination percentage was similar in both treated and untreated seeds.
Chemical compounds. Imbibition of dormant seeds in a solution containing a compound that
induces germination is another alternative to breaking physiological dormancy. An obvious and
effective choice is imbibition of seeds in a GA solution. Other compounds that have been used to
break dormancy are potassium nitrate, ethrel, butenolide (the active ingredient from smoke),
hydrogen peroxide, and anaesthetic compounds (e.g., acetone, ethanol).
Light
or surrounding vegetation. Many seeds, especially small seeds, require light for germination. One
of the most common and best studied examples is lettuce. This figure compares germination of
lettuce seeds from a photosensitive genotype in dark vs. light. In lettuce and other species, the light
mediated induction of germination in seeds is governed by phytochrome, which implies that red
light induces germination and far-red light inhibits it. It has been determined that Pfr, the farred
absorbing or activated form of phytochrome, induces germination by promoting GA synthesis.
In some species, continuous light may inhibit germination. Onion and leek are examples of species
where seed germination has been reported to be inhibited by continuous light. In these cases, seeds
must be germinated in the dark.
Leaching. Physiological dormancy may be overcome by modifying the balance of compounds that
inhibit and promote germination in the seed. In some cases such as beet seeds, dormancy may be
alleviated by exposing seeds to running water that dilutes or removes the inhibitory compounds from
the seeds. This treatment is known as leaching.
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Alternating temperatures. In addition to light, temperature fluctuation is another signal for a seed
regarding its relative position in the soil or surrounding vegetation. Deep in the soil and in the middle
of abundant vegetation, temperature fluctuations during the day and night are lower than near the
soil surface or in a gap without surrounding vegetation. This variation may explain, in part, why
seed germination in some species is favored by alternating temperatures compared to a constant
temperature; e.g., 30/20°C day/night versus a constant 25°C. Changes in hormone sensitivity have
been suggested as a possible mechanism mediating induction of germination by alternating
temperatures.
Priming treatments enhance germination percentage, uniformity and rate in several species.
Additionally, in some species such as lettuce, priming treatments alleviate thermoinhibition and the
light requirement for germination.
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2. Soil: Few prospective sites should be examined for both physical and chemical properties. For
this purpose profile pits of 2m depth should be dug in each representative part of the site as
suggested by external appearance, samples should be collected and analyzed for deciding the
choice. Soil samples must be analyzed to know the suitability of soil for growing fruit crops. Soil
analysis gives information on the type of soil, its fertility; its pH value etc. As far as possible flat
land should be selected. There should be no hard pan up to a depth of 2m.
3. Irrigation facilities: Most of the horticulture crops are raised under irrigation. So the water
facilities should also be taken in to consideration (quantity and quality).Water table should be
below 2 m depth.
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4. Nearness to the market: Saves the over head charges in transport and gives close touch with
market tastes (in the case of market gardens).In most cases a large percentage of the retail price
of fruits is accounted for by transport charges.
5. Transport facilities: Fruits being perishable cannot be moved for long distances without quick
and refrigerated transport. Bananas from the south are not reaching northern market in our own
country owing to the absence of refrigerated transport. But under refrigerated conditions, they can
be transported to longer distances. So; the orchards must be located where there is quick transport,
preferably a refrigerated transport system.
6. Power (electricity) supply: It would be a great advantage if electric power lines are running in
the proximity of the area as it can be tapped easily.
8. Availability of labour: Large orchards are started often in out of the way places and forest areas
away from populated centres.It would therefore be necessary to ensure that adequate labour is
available for orchard operatins.This point is of important in plantation crops particularly.
9. Social factors: These assume importance when large contingents of labour and managerial staff
are to be employed as plantations or large orchards. They should be provided with medical and
educational facilities, so that, they are content and stick on to the jobs.
(B) Fencing
Encroachment of trespassers, theft and animal grazing are the matter of common concern in fruit
orchard. Live fencing is an age old and traditional practice. The fences act as wind break also if
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Fundamentals of Horticulture
grown by using live plants particularly in the areas where cold and hot winds are the problem. The
main purpose of fencing is to protect the plants from stray animals and harmful effects of cold and
hot waves. Besides protection, the main function of live fences is to provide fuel wood, fodder and
food, and enriches the soil. Therefore, fencing the orchard is an important aspect in fruit cultivation.
Traditional farmers have vast knowledge about the practice of live fencing and the species used.
Planting live fence with tall growing trees as first row and small, bushy thorny plants in staggered
manner as second row along the boundary of orchard is very much important. In sloppy lands,
windbreaks should be planted along with contour in ascending and descending order to protect the
plants even at higher points. At such places, the live fencing cannot be combined with wind break
and must be planted along the boundary.
Fast growing hardy plants species can be grown as fence post at a spacing of 4-5m. These plants
commonly grown by seeds or maintained from natural vegetations growing along the boundary to
have strong root system. Farmers can also plant stakes of easy to root species such as Gliricidia
sepium, Moringa, Flame tree etc. The live fence posts are more durable than traditional wooden
poles as they are less prone to attack by termites and decay fungi. When grazing animals are part of
the farming system, the only way to establish live fence posts and a living fence, is to start with a
conventional wire fence supported by wooden pole and to gradually establish live fence plants to
substitute for the decaying poles. Gliricidia live posts fence can be established by planting a few
large (1.5-2.0 m) stakes in the existing conventional wire fence. These stakes normally take root
within a month or so. After the first pruning, subsequent pruning can be carried out every 4 to 8
months. Shoot pruning at intervals of 6 to 8 months result in woody sprouts that are suitable for use
as stakes. These pruned twigs can be used to multiply live stakes for fence posts within a year or
two after establishing the first live fence posts. Palmyra palm, Babool, Simaruba, Jamun, Jangal
jalebi, etc. can also be planted as fence posts.
Some hedge plants make a thicker and denser fence. Such plants include a number of species which
do not require additional support of barbed wire. To minimize the initial cost of orchard
establishment, it is advisable to establish dense and thorny hedges to protect the orchard. Another
alternative often used by farmers is the combination of easy to establish live fence plants that are
poisonous and unpalatable species. If well established, these natural barriers can deter both animal
and human trespassers from entering into the orchard. A live fence should ideally be planted just
before the monsoons and watered regularly after the rainy season is over to ensure optimum growth.
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Fundamentals of Horticulture
Agave, Sisal, Boradi, Karonda and Mehandi etc. are the promising plant species which can be
planted as live hedge along the orchard boundary.
(C) Windbreak
Tall trees are planted on the border of the orchard to reduce the impact of wind on the fruiting of the
fruit trees. Windbreak is commonly tall trees having dense foliage and keeps the surrounding
atmosphere humid. These trees help in minimizing the wind velocity, impact of high temperatures
which adversely affect the young plants and cause excessive fruit drop. Trees, which are generally
used as windbreak, are seedling mango, jamun, mulberry, moringa, jackfruit, carambola, shisham,
teak, gamhar and bamboo. These can also be used as live fencing posts. These trees are usually
planted close to each other to provide an effective shield. Tall growing trees like shisham and jamun
are planted 6m apart while low headed trees like mulberry and carambola are planted at 7m distance.
Normally, plants grown as wind break may extract nutrients from fruit plants. Therefore, a trench
of one meter depth should be dug between fruit plant row and wind break.
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4. Quincunx system:
This system is exactly like the square system but one additional tree is planted in the
centre of each square. The number of plants per acre by this system is almost doubled than
the square system. Fruit trees like papaya, kinnow, phalsa, guava, peach, plum etc. can be
planted as fillers in the permanent trees provides an additional income to the grower in the
early life of the orchard. The filler trees are uprooted when the main orchard trees start
commercial fruiting.
5. Triangular System; A pattern in which plants in alternate rows offset half the space between
plants in a row. The distance between the rows is the same or more than that in a row. Thus a
series of iso-scales triangles (two sides equal, instead of three as in an equilateral triangle) are
formed. This is easier to layout than the hexagonal pattern but results in 9% fewer plants than
the equivalent square or rectangle.
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Fundamentals of Horticulture
6. Contour system:
This system is usually followed in the hilly areas with high slopes but it is very much
similar to the square/rectangular system. Under such circumstances, the trees may be well
planted in lines following the contour of the soil with only a slight slope. Irrigation and
cultivation are then practiced only across the slope of the land as this practice reduces the
chances of soil erosion. In this system layout is done as in square/rectangular system, first by
establishing the base line at the lowest level and then marking for the trees should be done
from the base to the top. Bench terraces are used where the slope is greater than 10 per cent.
High density planting is one of the important methods to achieve higher productivity per unit area
both in short duration and perennial horticultural crops. Such system produces precocious cropping,
high and regular yields of good quality fruits and low labour requirement to meet ever rising
production costs.
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Fundamentals of Horticulture
Cluster Planting
Digging and filling of pits: Generally the pits are dug 2 to 3 months in advance of planting i.e.
March to May. Allow the pits to weather. A planting board (a plank about 1.5m long or longer
with two end notches and a center notch) is applied to the marking peg by its central notch and
two pegs are driven at the end notches. Then the board and the marking pegs are removed and a
pit of 1 -meter cube is dug. The two pegs driven at the end notches remain in position on either
side of the pit. All pits are dug similarly so that plant position is not altered at planting time.
While digging, the top soil should be kept on one side and th e bottom soil on another side
separately as the topsoil is somewhat fertile than the bottom soil. While filling the pits, the
topsoil is mixed with farmyard manure or compost, leaf mould or green leaf and a kilogram of
super phosphate. Then the pits are fi lled with the bottom layer of soil first and then with the
topsoil mixed with the manures. The soil after filling should rise about a foot over the orchard
level so as to allow for shrinkage on setting.
Filling is done a fortnight or two after digging pits . The pits are filled with a mixture of Top soil;
FYM, leaf mould and bone meal. Pits are filled a few inches
above the ground level for shrinkage and settlement.
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Characteristics of HDP
The trees of HDP should have maximum number of fruiting branches and minimum number
of structural branches.
These branches should be so arranged and trained in such a way that each branch casts a
minimum amount of shade on other branches
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Fundamentals of Horticulture
The height should be one and half its diameter at the base. A key to successful HDP depends
upon the control of tree size
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Fundamentals of Horticulture
Well trained trees are also much easier to prune than untrained tree.
Training is a broad term involves tying of branches to posts or tied or supported over a
trellis or pergola in certain fashion or sum of its parts are removed or trimmed with a view to give
the plant a particular desired shape.
Objectives of Training
The main objective of training is to control and regulate shape of trees so that orchard cultural
operations, harvesting can be done easily.
To admit adequate sunlight and air to the centre of the tree and to expose maximum leaf area
to the sun.
To build the strong frame work of tree and arrangement of scaffold branches.
To help to have a better crotch angle between scaffold branches of the tree.
To build the structure of the tree in such heights at which trees are less exposed for sunscald
and wind damage
Training develops a balance between vegetative and reproductive growth of tree.
To limit the growth and spread of the tree so that various cultural operations such as spraying
and harvesting are performed at minimum cost.
1. Central Leader:
This system of training is adopted in such types of trees which have a pronounced apical
dominance. Here the main trunk grows undisturbed. On account of vigorous and rapid growth of the
main trunk the tree develops a close center and grows to great heights. A central leader tree has one
main, upright trunk, referred to as the leader. Branching begins on the leader 75 to 100cm above the
soil surface to allow movement under the tree. In the first year, three to four branches, collectively
called a scaffold whorl, are selected. The selected scaffolds should be uniformly spaced around the
trunk, not directly across from or above another. Space scaffold whorls every 50 to 60 cm up the
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Fundamentals of Horticulture
central leader, leaving alternate areas without any branches to allow light into the centre of the tree
to the desired maximum tree height. The shape of a central leader tree is like that of a Christmas
tree. The lowest scaffold whorl branches will be the longest and the higher scaffold whorl branches
will be progressively shorter to allow maximum light penetration into the entire tree. Example, Pear
2. Open Center:
In this system, the main stem is allowed to grow only up to a certain height by heading
within a year of planting and all the subsequent vegetative growth promoted by lateral branches.
These results in a low head and as such the bulk of crop come closer to the ground. In contrast to
the central leader system, all sunshine is equally distributed to all the branches in open center
system. The open center trained trees are more fruitful resides greatly facilitate the operation like
spraying, thinning and harvesting. However, the branches form weak crotches since the branches
arise very close to one anothe r almost from the same point. So there is a certain amount of risk of
splitting of the branches when there are a heaver number of fruits on the trees. In areas of high
light intensity, such trees suffer from severe sunscald injuries, e.g Peach, Apple etc.
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Fundamentals of Horticulture
Methods of pruning
1. Thinning out: When a shoot is removed entirely from the inception (from the point of
origin) so that, no new shoot arises from that place, it is referred as thinning out. This thinning is
practiced in the removal of shoots arising in unwanted places, water shoots etc.
2. Heading back: When the branches grow tall and vigorously without producing flowers,
these shoots are headed back. When only one-third to one-half terminal portions of the branches,
having their basal portions intact, are removed, it is referred as Heading Back. When the branches
having their basal portions intact are removed the apical dominance of the twig, shoot or branch is
destroyed and the lateral buds are stimulated to growth. When a branch is cut almost to the base,
leaving a few inches of stump, carrying few buds. These buds left on the stump will give rise to
shoots which are important to the tree either being spur bearers or bearing flower buds or filling up
of gaps in the tree or forming vegetative wood from which flowers may arise in the following year.
The shoot from the bud nearest to the cut takes the place of the pruned shoot.
3. De-blossoming: Removal of surplus flowers to enable the tree to produce crops regularly
year after year is called deblossoming.This is practiced in alternate bearers like mango. apple etc.
Objectives of Pruning
To distribute the fruiting wood in all directions and to maintain a balance between
vegetative and reproductive phases.
To thin out branches so as to admit more light into the interior of the tree top so that the inner
wood also becomes fruitful.
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juvenility differs amongst plant species. The juvenility period of annual plants is quite brief, while
perennial plants have a long juvenility period.
Characteristics of Juvenility
Leaf Form- In its juvenile stage, the leaf is very active. Some plants, such as Acacia sp., have
different leaf morphologies in their juvenile stages.
Growth Form- The young branch resembles a whip. The branch develops in a straight line parallel
to the main stem. Citrus, Guava, and other fruits have such a branch, which is known as a water
sprout.
Presence of thorns- Some fruit trees have thorns when they are young or juvenile, but these thorns
vanish as they reach adulthood. Apple, pear, citrus, and other fruits are examples.
Leaf Retention In the juvenile stage, plants do not lose leaves all year. In the adult or mature stage,
however, it is shed.
Bud- The bud is an immature branch system that is frequently enveloped by scale leaves for
protection. A lateral branch, a flower, or an inflorescence emerges from the bud. The development
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Fundamentals of Horticulture
of these buds into flowers or inflorescences is an important part of fruit production. The phases of
flower growth are as follows:
1. Flower bud differentiation- Flower induction is another name for it. Bud differentiation requires
increased cell division below the apical section of the meristem in the middle area. Parenchyma cells
develop into flower primordia that surround the meristem as a result of cell division.
3. Development of Flowers- This includes the time between induction and bloom. The bloom
opening (anthesis).
A flower is a modified reproductive branch, which is essentially a stem with an apical
meristem that produces leaf primordia.
Types of buds
1. Simple Bud It grows from vegetative branches. Leaf bud is another name for it.
2. Mixed bud- It takes the form of a flower-bearing stalk. A flower bud is another name for it.
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Fundamentals of Horticulture
Hormonal balance
Change in the distribution of nutrients inside the apical meristem
Development of flower buds does not start unifromlyughout
thro the tree. In apple the
terminal bud of a spur begin their transformation
-6 weeks
4 earlier than lateral buds
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Fundamentals of Horticulture
In an orchard all the fruit trees do not bear equally or regularly and sometimes fail to flower and
fruit under similar conditions where another fruit tree bears heavily.
This failure to set fruits may be attributed to unfruitfulness.
To understand the problem of unfruitfulness in orchards a familiarity with following terms is
necessary.
1. Fruit setting: It refers to initial growth of ovary and its associated parts after blossoming and
taking it to maturity.
2. Fruitfulness: It is the state of plant when it is not only capable of flowering and fruit setting but
also takes these fruits to maturity and inability to do so is unfruitfulness or barrenness.
3. Infertility: Ability of a plant not only to produce fruits but develop viable seeds and the inability
to do so is referred as sterility or infertility. All fertile plants are fruitful but all fruitful plants are
not fertile (Seedless fruits).
4. Self fruitfulness: Ability of a plant to mature fruits after self pollination.
5. Self fertility: Capacity of a plant for the production of viable seeds after self pollination.
The ability of a plant to produce optimum crop is Fruitfulness. Where as, The
inability to achieve this is referred to as Unfruitfulness.
o This unfruitfulness is one of the serious problems of many orchards and its causes need to
be understood properly for effective control and obtaining economically acceptable production
level.
o The causes to this problem can be many and they have been broadly grouped into two
categories
(A) Internal factors
(B) External factors.
A. Internal factors associated with unfruitfulness: There are a number of internal factors which
are associated with unfruitfulness or sterility. They have further been categorized into three major
categories, they are
1. Evolutionary tendencies.
2. Genetic influence.
3. Physiological factors.
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Fundamentals of Horticulture
When stigmatic receptivity period does not coincides with pollen viability in monoecious plants it
is known as dichogamy.
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Fundamentals of Horticulture
In dichogamy self pollination is prevented in perfect flowered plants, due to maturity of two sex
elements at different times.
If the stamens ripe before the stigmas become receptive the flowers are known as protoandrous
and if stigmas become receptive before the stamens produce viable pollens it is known as
protogynous. This results in low production of fruits.
Protogyny is present in monoecious plants like walnuts, hazels, etc. whereas protandry is present in
many coconut varieties.
Majority of dioecious plants are also protogynous.
(iii) Abortive Flowers or aborted pistils or ovules:
2. GENETIC INFLUENCES
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Fundamentals of Horticulture
Self sterility is a condition determined by the inheritance received but can develop in favourable
Self sterility and self unfruitfulness has been reported in apple, pears, plums, almond, apricot, the
fails somewhere between production of functional gametes and the fusion of sex cells.
3. PHYSIOLOGICAL INFLUENCES:
(i) Slow pollen tube growth:
Slow growth of the pollen tube results in unfruitfulness.
Differences have been found in the rate of growth in selfed and crossed apples, pears, cherries and
certain citrus fruits.
This may be considered one type of incompatibility due to chemotropic or hormone influences.
Besides this, fertilization should take place within a short time failing which abscission will take
place at the base of the style, ovary pedicel or peduncle and fruit setting does not take place.
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Nutritional condition of spurs has positive correlation with fruit setting in apple. Spurs on vigorous
limbs with large leaves set more fruits than those borne on weak limbs.
More flowers ultimately lead to more fruit set and more flowers are generally borne on strong
limbs. Likewise flowers borne singly set fruits and mature as fruit and majority of those borne in
clusters drop down.
Ringing or girdling also lead to accumulation of an extra store of food material leads to fruitset and
develop parthenocarpically.
In the process of fruitifications the embryo is more important for development i.e. if nutritive
condition is favourable, it accompanies the development of the seed coat and fruit wall, if not, only
the latter portions are in high degree retardation in development.
Under insufficient nutrient supply the numbers of seed forming ovules are diminished and under
extreme nutrition deficiency both fruit wall and large number of ovules are diminished leading to
enabling to form seed.
In case of green house cucumbers, nutritional deficiency leads to arrest of growth of growing fruits
depending upon the position of the fruits and time of pollination. If a few of the cucumbers are
harvested remaining fruits resume growth.
In case of strawberries producing bisexual flowers may lead to produce pistillate flowers if
nutritional deficiency was observed.
However, nutritive condition has indirect influence on compatibility.
B. Unfruitfulness associated with external factors
1. Nutrient supply:
In certain families like Graminae, Cruciferae and Leguminaceae sterility normally occur due to over
feeding.
in poor soils.
High fertility level is generally associated with good pistil development and low level with poor
pistils and good stamens in grapes.
In olives low fertility leads to partial or complete degeneration of pistils.
2. Pruning and Training: Pruning tends to produce more true hermophrodite condition in grape
pistils.
3. Locality: Jonathan apple which is sterile in one location is reported to be self fertile in another
location.
4. Season:
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Fundamentals of Horticulture
Maintenance of critical nutrient status in tree l eaves for optimum crop production by adopting
correct nutritional programme based on plant and soil analysis.
In crops requiring regular pruning standard practices will have to be adopted based on crop,
variety and its phenology.
Unfruitfulness due to pathogens should be managed through effective plant protection
measures following integrated approach.
Problem of unfruitfulness due to tendency of alternate bearing should be over come through
replacement of regular bearing vari eties and crop regulation.
It is important to analyse the problem and then corrective measures should be suggested.
Basically the planning should be so done that future is problem free and then should be followed
by adoption of correct package of practice s.
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Fundamentals of Horticulture
pollination occurs, the pollen grain must germinate and grow into the ovary of the flower where the
male pollen cell unites with the female egg cell in a process called fertilization. The success of
pollination and fertilization depends upon the favorable environmental conditions as well as the
right pollen grain reaching the pistil of the right flower, as pollen can only fertilize specific,
compatible flowers. For example, pollen from a peach flower will not pollinate apple flowers
To better select fruit for planting, growers should become familiar with the terms used to describe
pollination characteristics and fruitfulness of different fruit types. Some of the most basic terms that
need to be understood are, self-pollination cross-pollination, self-fruitful, crossfruitful,
parthenocarpic, and perfect-flowered.
Self-pollination occurs when flowers are pollinated by pollen within the same horticultural variety
from the same or different trees. Most peach varieties, such as Red Haven, are fruitful when self
pollinated and therefore can be planted in very large blocks without using a second variety.
Cross-pollination occurs when flowers of one variety are pollinated by pollen from a second
variety. For example, Golden Delicious variety is often used in apple orchards to cross-pollinate
Red Delicious varieties.
Self-fruitful implies that a single variety of a given fruit type will produce satisfactory fruit crops
when grown by itself. This may occur because the variety is self-pollinating (such as peach) or
because they are parthenocarpic (such as some persimmons, figs, and satsumas).
Cross-fruitful implies that cross-pollination is required among two or more varieties to produce
satisfactory crops. Red Delicious apple varieties, for example, are cross-fruitful when
crosspollinated with variety of Golden Delicious.
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Fundamentals of Horticulture
Parthenocarpic basically means fruit are produced without complete seed development,
resulting in seedless fruits. Satsuma, for example, has sterile pollen, mostly nonviable ovules,
and is highly parthenocarpic, w hich results in the production of seedless fruit.
Perfect-flowered means that flowers of that variety have functional male and female parts.
Carlos is a perfect -flowered muscadine grape that is self-fruitful and is used as a pollinator for
female type varieties such as Fry.
Pollinator: Bees, wind, moths, butterflies, other insects, animals, and humans, may act to
physically transfer pollen from stamens to the stigma of the flower pistil.
Pollinizer: The source of pollen. Usually used when referring to cultivars planted for the purpose
of crosspollination.
Whether a fruit type is self -fruitful or requires cross -pollination influences how varieties are
arranged in a planting. For self -fruitful plants, single varieties perform well when planted alone.
For fruit types requiring cross pollina tion, two or more varieties of each type should be planted.
Planting entire rows with the same variety makes management of cultural practices and
harvesting much easier and more cost effective. When only the minimum number of pollinators
is desired, a pollinator variety should be planted as every third plant in every third row.
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Fundamentals of Horticulture
Plant bio-regulators (PBRs): It refers to both natural as well as synthetic compounds which affect
the growth and developmental process in plants. It includes plant hormones, synthetic and
physiologically active growth substances, growth inhibitors, synthetic enzymes, vitamins, organic
acid and other compounds which affect the biological activity of the plant system.
Plant growth regulators (PGRs): It refers to organic compounds other than nutrients which in
small quantities promote, inhibit or otherwise modify the physiological process of the plant.
Plant hormones or phytohormones: It refers to the compounds, produced within the plant system
which usually move within the plant from the site of production to the site of action and regulate
plant physiological process in physiologically very low concentration.
These phytohormones are classified into five broad groups- Auxin, Gibberellins, Cytokinins,
Ethylene and Abscisic acid. However, apart from these other phytohormones are brassinosteroides,
jasmonic acid, oligosaccharides, fusicoccin has also been reported as plant hormones.
1. Propagation: Most of the fruit crops are vegetatively propagated except papaya, capegooseberry,
phalsa, mangosteen. Sexual propagation is the only means for hybridization and rootstock production.
Poor seed germination in some of the fruit crop is major hindrance for sexual propagation. Gibberellins
(200-500 ppm) are mostly used for seed germination and substitution of chilling requirement.
Application of IBA @ 500-1250 ppm for soft wood, 1000- 3000ppm for semi hardwood and 2000-5000
ppm for hardwood is common for the initiation of rooting in cutting in different fruit crops. Apart from
auxin, a large number of physiologically and chemically unrelated compounds such as phenols and
allied simple aromatic compounds, glucosides, growth inhibiting, growth retarding and ethylene
producing chemicals significantly influence the rooting of cuttings. In a few cases, growth retardant
such as SADH has also proved beneficial in root induction. GA has antagonistic effect on rooting of
cutting.
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2. Use in micropropagation: Auxin and cytokinin are used to support basic level of growth of explants
grown under in vitro condition. For shoot initiation high cytokinin: auxin (BA: IAA) ratio is required
while for root induction high auxin: cytokinin (IBA: BA) is essential. Usually 0.1 and1.0 mg/l Benzyl
Adenine (BA) helps in terminal and lateral shoot formation in almond-peach hybrid under in vitro
condition, respectively while 0.1-0.2 mg/l IBA is most suitable for root induction.
Somatic embryogenesis from nucallar explants occur in the presence of 2,4-D at low concentration.
3. Breaking of seed and bud dormancy: Seeds of many fruit crops often require an extended period of
after ripening at low temperature before germination. Breaking of rest is of great significance to
overcome dormancy. GA (200-500 ppm) is used for accelerating seed germination in different fruit
crops. The treatment of GA (200 ppm) significantly reduced/substituted the period of seed stratification.
Termination of bud rest by GA (200-250 ppm) spray is a common practice in temperate fruit. It also
successfully reduces the period of chilling requirement.
4. Control of vigour: In fruit crops, tree size control is important for producing dwarf trees suitable for
high density orcharding. The use of different growth retardant like SADH/ paclobutrazol is effective in
reducing the growth of apple, apricot, lemon, litchi, mango, pear, peach, plum.
5. Flowering: Ethylene is the active principle responsible for flowering in pineapple. Apart from this
Acetylene, calcium carbide, ethephon, NAA are other chemicals which helps in floral induction in fruit
crops. NAA @ 10-15 ppm alone and Ethrel or ethepon @ 25 ppm in combination of urea (2%) and
CaCO3 (0.04%) is very effective for the induction of flowering in pineapple.
Paclobutrazol (PP333) has shown great promise for regulating flowering fruiting in mango. PP333
(800-1000 ppm) alone or in combination of urea (2%) successfully induced vegetative flush in
fruiting shoot of mango simultaneously.
NAA replaces girdling for improved flowering by mobilization of assimilates in the tree while
SADH promotes flowering in apple, pear, peach, lemon, blue berry and reduced shoot growth. Grape
and lemon respond to cycocel (CCC) treatment for increased flowering.
6. Parthenocarpic fruit development: In general, auxins (IAA, IBA, phenoxy acetic acid) -induce
parthenocarpy in fruit crop particularly in pineapple, citrus, banana etc. In apple, pear, peach, apricot,
almond- GA while in Grape- cytokinin and in Fig- 4-CPA is commonly used to induce parthenocarpy.
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7. Fruit thining: Auxins at higher concentratin (100-300 ppm). In grape- GA @ 40-60 ppm at postbloom
stage
8. Fruit growth and maturity: Auxins- increase fruit size. In grape - GA (40 -60 ppm) alone or in
combination with brassinosteroids after fruit set increases fruit sized and shape.
9. Control of fruit drop: Auxins (2,4 -D, NAA, 2,4,5 -T) at low concentration (10 -30 ppm) is very
effective for preventing fruit drop in different fruit crops
10. Improvement of fruit quality: In grape - GA (40 -60 ppm) at colour break stage increase TSS and
sugar content and reduced acidity.
11. Fruit ripening: Ethylene @ 250 ppm at fruit maturity is effective for ripening of fruits.Ethephone
@ 1000 ppm - causes degreening in citrus whileCycocle (500 ppm) applied twice at 15 days interval
at early maturity in Kagzi lime- results harvesting 20 day earlier.
12. Harvesting: Ethrel- in temperate fruit while cycloheximide - in mango, citrus is most effective for
this purpose. However, GA - 10 ppm inhibit degreening process and extending the shelf life of the
fruit
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Fundamentals of Horticulture
When to irrigate?
The time when a plant needs irrigation can only be judged by a keen observing eye.
The plants need water when their new leaves begin to show a wilting appearance. A little
before the trees show the sign of wilting.
The shedding of broad leaves in orchard shows distress symptoms.
successive irrigations.
fruiting of the trees while, over
irrigation serves no useful purpose and it may even prove to be harmful.
once. That may result in the splitting of fruits and even the splitting of bark of the branches and
trunk.
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SYSTEMS OF IRRIGATIONS:
Different systems of irrigation are followed in different parts of the country. The best system is the
one which meets the moisture seepage and evaporation.
c. Perforated pipelines.
III. Overhead or aerial irrigation : a. Sprinkler b. b. Revolving nozzles
IV. Drip or trickle irrigation
I. SURFACE IRRIGATION:
a. Flooding:
When the land is flat, letting in water from one end floods the entire area.
This system is commonly practiced in canal or tank bed areas.
It is the easiest method and permits the use of bullock drawn implements in the
orchards.
But in this there is wastage of water and leads to soil erosion also.
It encourages growth of weeds and spread of diseases like gummosis in citrus and
collar rot in papaya.
b. Basin system:
In this system, cir cular basins are provided around the trunk of the tree.
The basins are inter -connected in series and are fed through the main channel
running perpendicular to the tree rows.
When compared to flooding, this system minimises the loss of water.
In this system of irrigation, the water close to trunk may bring about certain
diseases like gummosis and nutrients are likely to be carried over from one
basin to the other.
c. Furrow system:
Unlike the flood system, here the entire land surface is not covered with irrigation water.
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Fundamentals of Horticulture
the trees.
Water is let in these furrows from the main channels.
In orchards, two furrows on each side of the rows are generally made.
It is suited to such lands, which have a moderate slope to the extent of 1 -2% if
the water is to run freely and reach the ends of the furrows.
Where the slope is sharp, the furrows are made to follow the contour more or
less closely.
This method has disadvantage of excess of water penetration at the head than
at the farther end, which may result in variation in vigour and growth of trees.
d. Ring system:
This is an improvement over the basin system.
In this system, a ring is formed close and around the tree and water is let into the
basin.
This method is recommended for citrus trees thereby reducing the chances of
collar rot to which these trees are often susceptible.
The size of the ring will increase as the tree grows.
In this system, the spread of diseases like collar rot, etc., are prevented.
However, it involves more labour and capital and it does not permit uniform
distribution of water throughout the bed or basin as in the basin system of
irrigation.
SUB-SURFACE IRRIGATIO N:
In general, however, it must be used with great caution because of the danger of water
logging and salt accumulation.
If the sub-strata are so slowly permeable that practically no water moves through, water
added may stand in soil sufficiently for long time which results an injury to the plant root due
to poor aeration.
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Fundamentals of Horticulture
Where irrigation water or the sub-soil contains appreciable amount of salt, sub-soil
irrigation is usually not advisable.
Land must be carefully levelled for successful subsoil irrigation so that raising the water
table will wet all parts of the field equally.
May have definite economic advantages in developing new land that has never been
irrigated, particularly where the land is rough or the soil is too much porous, shallow or
highly erodable.
It is quite useful where only small streams are available, such as irrigation wells of small
capacity.
It is helpful in irrigatin g at the seedling stage when the furrowing is difficult and flooding
leads to crusting of soil.
Fertilizer materials may be evenly applied by this method.
This is usually done by drawing liquid fertilizer solutions slowly into the pipe.
It has several disadvantages like
High initial cost,
Difficult to work in windy location,
Trouble from clogging of nozzle,
Interference in pollination process and
Requirement of more labours while removing or resetting.
In general, this system is best adopted for areas where ordinary surface systems are
inefficient.
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Fundamentals of Horticulture
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Fundamentals of Horticulture
The Nutrients are chemical elements which are absorbed by the plants in more or less quantity to
transform light energy into chemical energy and to keep up plant metabolism for the synthesis of
organic materials.
These materials constitute among other things, foods for humans and animals and a range of raw
materials for various industrial uses.
Feeding of plants with nutrients is termed as nutrition.
Successful growth and production of the plants in general requires a proper supply of the 16
elements. These elements are regarded as essential to life in higher plants.
Allen and Arnon (1955) laid out following criteria for categorising nutrients essentiality to plants:
1. Complete or partial lack of the element in question must make normal plant growth
impossible
2. Deficiency symptoms must be reversibly by the addition of the elements in question 3.
The element must play specific role in the plant metabolic symptom They are:
1. Basic elements : Carbon (C),Hydrogen(H) and Oxygen( O) (03)
2. Macro elements : Nitrogen ( N), Phosphorus( P) Potash( K), Calcium( Ca)
Magnesium (Mg) and Sulphur(S) (06)
3. Micro elements : Manganese (Mn), Molybdenum ( Mo), Chlorine (Cl), Zinc ( Zn),
Boron (B), Copper (Cu) and Iron (Fe) (07) Macro elements: The nutrients that are required in
relatively large quantity are termed as macro elements. Micro elements: are those required in
relatively less quantity are termed as micro nutrients.
Besides some nutrients like Aluminium (Al), Cobalt (Co), Sodium (Na), Silica (Si) and Vanadium
(V) are not considered necessary always because either their essential character has been proved
only in some plants or in certain metabolic processes that are not always necessary.
TYPES OF FERTILIZERS
Inorganic fertilizers
Industrially manufactured chemicals.
Contains higher nutrient than organic manures.
Nutrient input is lost through leaching, runoff, volatilization, fixation by soil or consumption by
weeds etc.
Organic fertilizers
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Fundamentals of Horticulture
These are plant and animal wastes that are used as nutrients after decomposition.
Improves the soil tilth, aeration, water holding capacity and activity of micro-organism.
WHERE TO APPLY THE MANURES?
In fully grown trees, the manures and fertilizers should be given over the area, where their
buds.
Disadvantages:
Some of the elements like phosphorous and potash do not readily move in the soil. Therefore,
surface application ma y not be available to the trees especially in drier tracks.
Leads to accumulation of potassium in surface soil beyond detrimental levels causing injury to
plants.
Surface application always stimulates weed growth.
Band placement:
Application of fertilizer on the sides of rows.
Fertilizer in solid and liquid forms can be applied.
Quantity of fertilizer may be economised.
RING PLACEMENT:
Commonly followed in fruit trees.
Fertilizers are applied in a ring encircling the trunk of the trees extending the entire canopy.
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Fundamentals of Horticulture
e to the leaves.
STARTER SOLUTION:
starter solution.
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Fundamentals of Horticulture
FEEDING NEEDLES:
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Fundamentals of Horticulture
But after the initial success, it had shown the symptoms of fatigue evident from the undesirable side
effects on natural resources, such as soil, water and biodiversity and thus human health.
The vast areas of soils once fertilizer was degraded due to soil erosion, salinisation or general loss
of soil fertility.
Water resources have been over-exploited and polluted due to excessive requirement of irrigation
water for high yielding varieties and intensive use of agro-chemicals.
Many plants and animal species were wiped out and are endangered.
Residues of harmful pesticide in food and drinking water endangered both farmers and consumer
health point of view and thus excessive use of external inputs consumes a lot of energy from non-
renewable resources.
Organic farming is a way of conserving the soil and maintaining the fertility, protect soil flora and
fauna/diversity.
It has lesser effect on pollution either of ground water, lakes and rivers.
Organic agriculture does not utilize non-renewable external input and energy.
Since no chemical or pesticide is used in crop production, there is very low chance of pesticide
residues in food.
At the same time the organic products are healthier and have better product quality like taste, aroma
and storability.
Input cost is drastically reduced in organic cultivation but the market price leading to higher income
for farmers.
Aims of organic production and processing:
To produce sufficient quantities of high quality food, fibre and other products.
To work compatibly with natural cycles and living systems through the soil, plants and animals in
the entire production system.
To recognise the wider social and ecological impact of and within the organic production and
processing systems.
To maintain and increase long-term fertility and biological activity of soils using locally adopted
cultural, biological and mechanical methods as opposed to reliance on chemical inputs. To maintain
and encourage agricultural and natural biodiversity on the farm and surroundings through the use of
sustainable production systems and protection of plant and wildlife habitats.
To maintain and conserve genetic diversity through attention to on-farm management of genetic
resources.
To promote the responsible use and conservation of water and all life therein.
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Fundamentals of Horticulture
To use, as far as possible, renewable resources in production and processing systems and
avoid pollution and wastes.
To foster local and regional production and distribution.
To create a harmonious balance between crop production and animal husbandry.
To provide living conditions that allows animals to express the basic asp ects of their innate
behaviour.
To utilise biodegradable, recyclable and cycled packaging materials.
To provide everyone involved in organic farming and processing with a quality of life that
satisfies their basic needs within a safe, secure and healthy working environment.
To support the establishment of an entire production, processing and distribu tion chain which
is both socially and ecologically responsible.
To recognise the importance of, and protect and learn from, indigenous knowledge and
traditional farming systems.
Organic food products exported from India:
Organic cereals: Wheat, Rice and Maize or Corn.
Pulses: Red gram and Black gram.
Fruits: Banana, Mango, Orange, Pineapple, Passion fruits, Cashew nut and Walnut.
Oilseeds and oils: Soybean, Sunflower, Mustard, Cotton seed, Groundnut and Castor.
Vegetables: Brinjal, Garlic, Potato, Tomato and Onion.
Herbs and spices: Chilli, Peppermint, Cardamom, Turmeric, Black pepper, White pepper,
Amla, Tamarind, Ginger, Vanilla, Cloves, Cinnamon, Nutmeg and Mace.
Others: Jiggery, Sugar, Tea, Coffee, Cotton and Textiles.
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