Prokaryotic Metabolism
Prokaryotic Metabolism
Metabolism
Prokaryote metabolism
• How prokaryotes get energy and nutrients. Chemotrophs and
phototrophs. Heterotrophs and autotrophs.
• Key points:
• Some prokaryotes are phototrophs, getting energy from the sun.
Others are chemotrophs, getting energy from chemical compounds.
• Some prokaryotes are autotrophs, fixing carbon from CO2.
• Others are heterotrophs, getting carbon from organic compounds of
other organisms.
Prokaryote metabolism
• Prokaryotes may perform aerobic (oxygen-requiring)
or anaerobic (non-oxygen-based) metabolism, and some can switch
between these modes.
• Some prokaryotes have special enzymes and pathways that let them
metabolize nitrogen- or sulfur-containing compounds.
• Prokaryotes play key roles in the cycling of nutrients through
ecosystems.
• Prokaryotes (bacteria and archaea) are way
more diverse than humans in their
nutritional strategies – that is, the ways
they obtain fixed carbon (fuel molecules)
and energy. Some species consume organic
material like dead plants and animals.
Others live off of inorganic compounds in
Introduction rocks. One bacterium, Thiobacillus
concretivorans, consumes metal-melting
sulfuric acid!
• In this Presentation, we will take a closer
look at the many ways that prokaryotes
obtain and metabolize food, and how they
can influence cycling of nutrients.
• All of Earth’s life forms need
energy and fixed carbon (carbon
incorporated into organic
molecules) to build the
macromolecules that make up
Nutritional their cells. This applies to
humans, plants, fungi, and, of
modes course, prokaryotes. Living
organisms can be categorized by
how they obtain energy and
carbon.
• First, we can categorize organisms by where
they get fixed (usable) carbon:
• Organisms that fix carbon from carbon dioxide
or other inorganic compounds are
called autotrophs.
• Organisms that get fixed carbon from organic
Nutritional compounds made by other organisms (by
eating the organisms or their by-products) are
modes called heterotrophs.
• In addition, we can categorize organisms by
where they get energy:
• Organisms that use the light (mainly the sun)
as a source of energy are called phototrophs.
• Organisms that use chemicals as a source of
energy are called chemotrophs.
We can divide prokaryotes •We tend to be pretty familiar with
(and other organisms) into photoautotrophs, such as plants, and
chemoheterotrophs, such as humans and other
four different categories animals. Prokaryote species fall into these two
categories, as well as the two less familiar
based on their energy and categories (photoheterotrophs and
chemoautotrophs) to which plants and animals
• Nitrogen-metabolizing
prokaryotes include nitrogen
fixers, nitrifiers, and denitrifiers.
They play key roles in the
nitrogen cycle by converting
nitrogen compounds from one
chemical form to another.
Nitrogen metabolism
• Nitrogen-fixing prokaryotes convert (“fix”)
atmospheric nitrogen N2 into ammonia NH3,
which plants and other organisms can incorporate
into organic molecules. Some plant species in the
legume family, such as peas, form mutually
beneficial relationships (mutualisms) with
nitrogen-fixing bacteria. The plants house and feed
the bacteria in structures called root nodules, and
the bacteria provide fixed nitrogen to the roots.
• Other prokaryotes in the soil, called nitrifying
bacteria, convert the ammonia into other types of
compounds (nitrates and nitrites), which may also
be absorbed by plants. Denitrifying
prokaryotes do more or less the reverse, turning
nitrates into N2gas.
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