Chap-06_Anatomy of Flowering Plants (27)_E
Chap-06_Anatomy of Flowering Plants (27)_E
Chap-06_Anatomy of Flowering Plants (27)_E
Syllabus
Morphology and modifications; Tissues; Anatomy and functions of different parts of flowering
plants: Root, stem, leaf, inflorescence- cymose and recemose, flower, fruit and seed (To be
dealt along with the relevant practical of the Practical Syllabus).
Chapter Index
THE TISSUES
The term tissue was coined by Grew.
A group of similar or dissimilar cells that perform a common function and have a common
origin is called tissue.
The tissues are classified into two main groups, namely, meristematic and permanent tissues
based on whether the cells being formed are capable of dividing or not.
A. Meristematic Tissues
These tissues consist of cells that retain the power of division.
The protoplasm within the cell is dense, the vacuole is smaller or absent.
These cells are isodiametric, without intercellular spaces.
The nucleus is bigger in size.
These cells have thin cellulosic cell wall.
Metabolically active cells with high surface area per unit volume and nucleo-cytoplasmic ratio.
Ergastic substances are absent.
Colourless proplastids are present in cells.
Concept Builder
Many theories have been given to explain shoot apex organisation, such as
(i) Apical cell theory. It was proposed by Hofmeister and Nageli. According to this theory a
single apical cell leads to the development of entire plant body. This theory is applicable to
algae, as well as to most of the bryophytes and pteridophytes.
Concept Builder
Regarding the organisation of root apex, following theories have been put forward.
(i) Korper-Kappe theory. It was proposed by Schuepp (1917). This theory is comparable with
the tunica and corpus theory of shoot apex because it is also based upon plane of division.
Korper means body and Kappe means cap.
Conceptual Questions
The meristem found intercalated between permanent tissues is called.
Give one word for the meristem that is required for increase in girth of plant body.
Ergastic substances are absent in meristem cells (True / False).
Secondary meristem arises directly from primary meristem (True / False).
Inverted cup like structure in root apex is called _ .
B. Permanent Tissues
They are composed of living or dead cells which are derived from the meristematic tissue but
have lost their ability to divide.
They are primary permanent tissues, if they are derived from apical or intercalary meristem.
They are secondary permanent tissues if they are derived from lateral meristem.
Permanent tissues are mature cells with permanent special structure and function.
These are of three types:
(I) Simple Tissues
(II) Complex Tissues and
(III) Secretory Tissues
(I) Simple tissues
They are made up of one kind of cells performing similar function.
(a) Parenchyma
These cells are found almost in all parts of plants such as roots, stem, leaves, fruits and seeds.
These cells are isodiametric, spherical, oval or polygonal with intercellular spaces.
These cells are living with thin cellulosic cell wall.
Elongated parenchyma with tapering ends is called prosenchyma.
Functions:
(i) The main function is storage of food , e.g., fruits and endosperm.
(ii) Storage of water in fleshy stem and leaves e.g., Opuntia.
(iii) Sometimes, they store secretory substances (ergastic substances) such as tannins, resins and
gums and they are called as idioblasts.
(iv) In hydrophytes, they have large intercellular spaces filled with air and are called aerenchyma.
They help in circulation of air as well as provide buoyancy to plants.
Concept Builder
Collenchyma is of three types :
1. Angular collenchyma - Angular walls thickened e.g., stem of Marigold, Tomato, Datura.
2. Lamellate collenchyma - Tangential walls thickened e.g., stem of Sunflower.
3. Lacunate collenchyma - Lacunate thickening, intercellular spaces are present e.g., stem of
Calotropis.
Function :
It provide mechanical support, flexibility and elasticity to the organs and due to peripheral
position in stems they resist bending and pulling action of wind . It is especially useful for
young plants and herbaceous organs where it is an important supporting tissue.
(ii) Vessels
They are also elongated and tube like, formed from a row of cells placed end to end.
The partition walls are either perforated or disappear altogether, resulting in an elongated tube.
Walls are thickened and lignified, may have annular, spiral, reticulate or scalariform
thickenings.
Vessels are dead at maturity and without nuclei.
The main function is conduction of water.
Vessels are advanced type of conducting elements.
Concept Builder
In pteridophytes and gymnosperms, vessels are absent (non porous wood).
Sometimes, primitive vessels are present in Gnetum and Ephedra (Gnetales).
Vessels are characteristic of angiosperms (porous wood), but they are absent in members of
vesselless families like Winteraceae, Trochodendraceae and Tetracentraceae.
Concept Builder
When parenchyma is diffused or not associated with vessels, it is called apotracheal
parenchyma and when parenchyma surrounds or is associated with vessels, it is called
paratracheal parenchyma.
At the end of growing season a callose plug (made of callose carbohydrate) is deposited on
sieve plate in old sieve tubes which inhibits the activity of sieve tubes.
In spring season the callose plug gets dissolved.
Self Assessment
Intercalary meristem is related to all, except
(1) Present between permanent cells
(2) Part of apical meristem
(3) Increasing the girth of axis
(4) Regenerates part of grasses removed by grazing herbivores
Find odd one out W.r.t. histogens found in shoot apex according to Hanstein
(1) Tunica (2) Periblem (3) Plerome (4) Dermatogen
Thickening material in collenchyma is
(1) Pectin, cellulose, hemicellulose (2) Lignin, cellulose, hemicellulose
(3) Hemicellulose, suberin, cellulose (4) Suberin, pectin, cellulose
Q.4 Long hair like sclereids found in aerial roots of Monstera are known as
(1) Brachysclereids (2) Trichosclereids (3) Macrosclereids (4) Astrosclereids
Q.5 Interxylary phloem is not found in
(1) Potato (2) Leptadaenia (3) Salvadora (4) Chenopodium
Q.6 Which component is not found in phloem of angiosperms?
(1) Albuminous cells (2) Sieve tube (3) Companion cell (4) Bast fibre
Q.7 Major xylary element in wood of a gymnospermic plant is
(1) Vessel (2) Tracheid (3) Xylem fibre (4) Xylem parenchyma
Q.8 Which of the following cell helps in maintaining the pressure gradient in the sieve tubes?
(1) Phloem parenchyma (2) Bast fibre
(3) Companion cell (4) Wood fibre
Q.9 Select an incorrect match
(1) Tunica – Corpus theory –Schmidt (2) Histogen theory – Hanstein
(3) Quiescent centre theory – Clowes (4) Korper – Kappe theory – Hofmeister
Q.10 Sclerenchyma fibres are absent in :
(1) Protoxylem (2) Protophloem
(3) Metaxylem (4) More than one option is correct
Ans. Q.1 (3), Q.2 (1), Q.3 (1), Q.4 (2), Q.5 (1), Q.6 (1), Q.7 (2), Q.8 (3), Q.9 (4), Q.10 (4)
Concept Builder
These tissues are of two types :
(1) Laticiferous tissues.
They are made up of thin walled, elongated, branched and multinucleate (coenocytic)
structures that contain colourless, milky or yellow coloured heterogenous substance called
latex.
These are irregularly distributed in the mass of parenchymatous cells.
These tissues are of two types –
(a) Latex cells. They do not fuse and do not form network. Such tissues are called simple or non-
articulated laticifers e.g., Calotropis (Asclepiadaceae), Nerium, Vinca (Apocyanaceae),
Euphorbia (Euphorbiaceae), Ficus (Moraceae).
(b) Latex vessels. They are formed due to fusion of cells and form network like structure. Such
tissues are called compound or articulated laticifers, e.g., Argemone Papaver (Papaveraceae),
Sonchus (Compositae), Hevea (rubber plant), Manihot (Euphorbiaceae).
The latex of some plants is of great commercial importance such as
(i) Source of commercial rubber is latex of Hevea brasiliensis, Ficus elastica, Cryptostegia,
Manihot glaziovii.
(ii) Source of chewing or chickle gum is latex of Achras sapota.
(iii) Source of enzyme papain is latex of Carica papaya. .
(iv) Source of alkaloid opium is latex from immature capsules of Papaver somniferum (Poppy) .
(2) Glandular tissues.
They include different types of glands which secrete oils, gums, mucilage, tannins and resins.
They may be :
(a) External glands. They generally occur on the epidermis of stem and leaves e.g., glandular hair
in Utricularia, nectar secreting glands in flowers, digestive enzyme secreting glands) in
Drosera, Nepenthes (insectivorous plants).
(b) Internal glands. These are present internally, e.g., oil glands in Citrus and Eucalyptus,
resinous ducts in Pinus (schizogenous origin) and mucilage secreting glands in leaves, of Piper
betel. The glands which secrete essential oils are called osmophores.
Stomata: A. With bean-shaped guard cells B. With dumb-bell shaped guard cells
Mostly epidermis is single layered parenchymatous, but is multilayered in leaf of Ficus and
Nerium.
Epidermis is mainly protective in nature (external protective tissue).
In grass leaves, motor or bulliform cells are present in upper epidermis.
On stem, the epidermal hairs are called trichomes, which are usually multicellular.
They may be branched or unbranched and soft or stiff. They may even be secretory. These help
in preventing water loss due to transpiration.
Concept Builder
(b) Pericycle. It lies between endodermis and vascular tissue. It is mostly single layered and
parenchymatous in roots and sclerenchymatous (mixed with parenchyma) in stem. The
pericycle cells just opposite the protoxylem are the seat for the origin of lateral roots. In
dicot roots, pericycle form a part of cambium and whole of cork cambium.
(c) Pith. It occupies the central part in dicot stem and monocot root. It is mostly made up of
parenchymatous cells. In dicot root, pith is completely crushed by the metaxylem elements. In
dicot stem the pith cells between the vascular bundles become radially elongated and are
known as primary medullary rays or pith rays. They help in lateral translocation.
Concept Builder
Concentric. When one vascular tissue surrounds the other. They are of two types:
(i) Amphicribal or Hadrocentric. The xylem is surrounded on all sides by phloem e.g., ferns.
(ii) Amphivasal or Leptocentric. The phloem is surrounded on all sides by xylem e.g., Yucca,
Dracaena.
Conceptual Questions
Source of enzyme papain is latex of .
The epidermal cells containing cystoliths are called .
Main function of general cortex is storage of food (True/False).
Name the type of vascular bundles, when the xylem and phloem are arranged on different radii,
alternating with each other.
Bicollateral vascular bundles are present in the stem of sunflower (True/False).
Anatomy of Stem
1. Primary structure of dicot stem
Dicot stem consists of following layers:
(i) Epidermis: It is the outermost layer consisting of single layer of closely arranged cells with
cuticle (cutinized). It bears multicellular hairs.
(ii) Cortex: It is differentiated into hypodermis, general cortex and endodermis. Hypodermis is
collenchymatous. General cortex is parenchymatous. Endodermis is wavy. It has starch grains
hence it is called starch sheath or endodermoid.
(iii) Pericycle : It lies inner to endodermis. Pericycle is few layered thick. Above vascular bundle,
it is sclerenchymatous and outside medullary rays it is parenchymatous.
(iv) Vascular bundles: These are in the form of a ring or eustele. They are conjoint, collateral
and open. In the family Cucurbitaceae, the stem is wavy, having five ridges and five furrows
and vascular bundles are present in ridges and furrows. Vascular bundles are bicollateral and
open.
Part of transverse sections of stem: A-B. Dicot stem C-D. Monocot stem
Anatomy of Leaf
1. Structure of dorsiventral leaf (dicot) :
In cross section of dicot leaf, following parts can be observed
Self Assessment
Q.11 Latex vessels are found in
(1) Ficus (2) Vinca (3) Euphorbia (4) Hevea
Q.12 Epidermal tissue system is derived in dicot root from
(1) Calyptrogen (2) Dermatogen (3) Periblem (4) Plerome
Q.13 Multilayered epidermis is found in the leaves of
(1) Nerium and Ficus (2) Ficus and Vanda
(3) Equisteum and Grasses (4) Vanda and Nerium
Q.14 Atactostele is found in the stem of
(1) Mango (2) Maize (3) Capsella (4) Sunflower
Q.15 Carsparian strips are found on radial and inner walls of
(1) Stem endodermis (2) Root endodermis (3) Pericycle (4) Outer cortex
Q.16 Dicot root is similar in all given characters with monocot root, except
(1) Radial, exarch vascular bundles (2) Unicelled root hairs
(3) Pericycle forms the lateral roots (4) Well developed pith
Q.17 Stem of barley is related to
(1) Presence of collenchyma in hypodermis (2) Scattered vascular bundles
(3) Presence of parenchymatous pericycie (4) Presence of wedge shaped vascular bundles
Q.18 a. Palisade tissue is present towards upper epidermis in monocot leaves.
b. Lower layer of mesophyll cells is loosely packed with few chloroplasts in dicot leaves.
c. Dicot leaves have conjoint, collateral and closed vascular bundles
(1) All are incorrect (2) Only B is incorrect
(3) Only A is incorrect (4) Only C is incorrect
Ans. Q.11 (4), Q.12 (2), Q.13 (1), Q.14 (2), Q.15 (2), Q.16 (4), Q.17 (2), Q.18 (3), Q.19 (2),
Q.20 (4)
SECONDARY GROWTH
Increase in girth or thickness or diameter of the axis occurs due to formation of new tissues as
a result of joint activity of vascular cambium and cork cambium in steler and extrasteler region
respectively.
It occurs in the root and stem of gymnosperms and dicots.
Secondary growth in dicot stem completes in following steps :
A. Formation of vascular cambium ring :
(i) Intrafascicular cambium : It is primary in origin and is present between primary phloem and
primary xylem.
(ii) Interfascicular cambium : It is true secondary meristem. It originates from parenchyma cells
of medullary rays region. It lies in between the vascular bundles.
(iii) Vascular cambium ring: Both intrafascicular and interfascicular cambia join together and
form a cambium ring.
Cells of cambium are of two types:
(a) Fusiform initials : They form tracheids, vessels, fibres and axial parenchyma in secondary
xylem and sieve tubes, companion cells, fibres and axial parenchyma in secondary phloem.
(b) Ray initials : These are isodiametric and form ray parenchyma (vascular rays).
(iv) Periclinal division in cells of vascular cambium ring.
(v) Formation of secondary phloem (outside the vascular cambium) and secondary xylem (inner to
vascular cambium). The amount of secondary xylem produced is 8-10 times greater than
secondary phloem.
(vi) Due to formation of secondary phloem primary phloem is crushed to death, known as
obliteration. Primary xylem being dead and lignified, persists in the pith region by replacing
the pith cells.
(vii) At some places, the cambium forms a narrow band of parenchyma, which passes through the
secondary xylem and the secondary phloem in the radial directions. These are the secondary
medullary rays.
(viii) Formation of secondary structures i.e. annual rings, sapwood and heart wood, hardwood and
softwood etc.
Concept Builder
Growth rings are distinct or sharply demarcated in the plants of temperate climate e.g.,
Shimla, Nainital, Mussourie due to presence of contrasting seasonal variations.
Growth rings are not distinct or sharply demarcated in the trees of tropical climate (near
equator) e.g., Calcutta, Bombay, Madras due to absence of contrasting seasonal variations.
Concept Builder
Wood of dicot trees is called porous or hard wood because it consists of vessels (pores).
The wood of gymnosperms does not contain vessels (pores) and is known as soft or non
porous wood.
Such wood consists of 90 to 95% tracheids and 5 to 10% of ray cells.
Sap wood will decay faster if exposed freely to the air.
On the basis of distribution and size of vessels, porous wood is of two types:
(a) Diffuse porous wood (primitive) : Vessels of same size are uniformly distributed throughout
the growth, e.g., Pyrus, Betula.
(b) Ring porous wood (advanced) : Large vessels are formed in early wood when the need of
water is great and small vessels are formed in late wood, e.g., Quercus, Morus.
Concept Builder
(i) Scaly bark: Develops in strips e.g., Acacia, Psidium.
(ii) Ring bark: Develops in the form of sheet or ring, e.g., Betula (Bhojpatra).
The outermost layer of bark is dead and called as rhytidome.
The bark of Betula was being used as substitute of paper in ancient time to write manuscript.
Conceptual Questions
Balloon like structures in vessels of heart wood are called .
Complimentry cells are cut off by phellogen towards outside (True / False).
Meristematic layer that is responsible for cutting off xylem and phloem is called vascular
cambium (True/False).
Self Assessment
Q.21 Vascular cambium in dicot stem is
(1) Primary meristem in origin
(2) Secondary meristem in origin
(3) Promeristem in origin
(4) Both primary and secondary meristem in origin
Ans. Q.21 (4), Q.22 (3), Q.23 (4), Q.24 (1), Q.25 (4)
Concept Builder
(i) Dicots with scattered vascular bundles in stem, e.g., Podophyllum, Peperomia, Piper, Papaver.
(ii) Cortical vascular bundles, e.g., Nyctanthus, Kalanchoe, Casuarina.
(iii) Medullary bundles, e.g.; Mirabilis, Bougainvillea, Amaranthus, Achyranthus.
(iv) Polystelic condition, e.g., Primula, Dianthera
(v) Anomalous or abnormal secondary growth occurs in Bougainvillea, Boerhaavia,
Chenopodium, Aristolochia.
(vi) Some monocots show abnormal secondary growth by meristematic tissue which develops
around vascular bundles, e,g., Dracaena, Yucca, Agave .
(vii) Virgin cork is first formed periderm.
(viii) Wound cork: It is secondary meristem; formed below injured area. It forms cork on outer side
and callus below; which heals the wound.
(ix) Abnormal secondary growth in dicot root occurs in Beet root (Beta vulgaris) and Sweet potato
(Ipomoea batatas) by the formation of numerous accessory rings of cambium which cut more
storage parenchyma in secondary phloem and less secondary xylem.
(x) Homoxylous wood-wood of vesselless dicots, e.g., Ranales (Winteraceae, Tetracentraceae,
Trochodendraceae) .
(xi) Heteroxylous wood is wood of vessel bearing dicots.
(xii) Polyderm is a special type of protective tissue occurs in roots and underground stems of
members of Rosaceae and Myrtaceae. Its outermost layer is dead and suberized.
Section–B
All given tissues are formed as a result of redifferentiation process, except
(1) Phellem (2) Phelloderm (3) Secondary xylem (4) Interfascicular cambium
A. According to Clowes root apex consists of an inverted cup like structure.
B. Low amount of RNA, DNA and protein is characteristic to waiting meristem.
C. Cells of reserve meristem can divide only when the root apex gets injured.
(1) All are correct (2) A & C are incorrect
(3) A & B are incorrect (4) B & C are incorrect
Isodiametric sclereids, found in hard endocarp of coconut and fleshy portion of some fruits are
(1) Brachysclereids (2) Asterosclereids (3) Osteosclereids (4) Trichosclereids
Members of winteraceae, tetracentraceae and trochodendraceae
(1) Do not have tracheids (2) Do not have albuminous cells
(3) Do not have vessels (4) More than one option is correct
Find set of cells connected by pit fields between their common longitudinal walls
(1) Companion cell and pholem fibres (2) Companion cell and sieve tube
(3) Sieve cell and albuminous cell (4) Sieve tube and pholem fibre
Seat of origin of late.ral root and formation of cork cambium are features related to
(1) Endodermis (2) Pericycle (3) Hypodermis (4) Pith rays
Answer Key
Section–A
Q.1 3 Q.2 2 Q.3 4 Q.4 3 Q.5 3 Q.6 4 Q.7 3
Q.8 1 Q.9 1 Q.10 2 Q.11 4 Q.12 2 Q.13 1 Q.14 2
Q.15 4 Q.16 2 Q.17 4 Q.18 3 Q.19 4 Q.20 4 Q.21 3
Q.22 2 Q.23 2 Q.24 1 Q.25 2 Q.26 1 Q.27 1 Q.28 4
Q.29 1 Q.30 2 Q.31 2 Q.32 1 Q.33 2 Q.34 3 Q.35 4
Q.36 4 Q.37 2 Q.38 2 Q.39 4 Q.40 2 Q.41 4 Q.42 1
Q.43 1 Q.44 2 Q.45 2 Q.46 4 Q.47 3 Q.48 1 Q.49 2
Q.50 3 Q.51 1 Q.52 4 Q.53 2 Q.54 2 Q.55 2 Q.56 2
Q.57 2 Q.58 2 Q.59 3 Q.60 4 Q.61 4 Q.62 3 Q.63 3
Q.64 4 Q.65 3 Q.66 4 Q.67 4 Q.68 1 Q.69 2 Q.70 1
Q.71 1 Q.72 1 Q.73 3 Q.74 2 Q.75 2 Q.76 1 Q.77 2
Q.78 3
Answer Key
Answer Key