Cell Wall, Cell Membrane
Cell Wall, Cell Membrane
Cell Wall, Cell Membrane
• Cell wall is a rigid, freely permeable and protective covering of plant cells (made up of
cellulose), fungal cells (made up of chitin) and prokaryotic cells (made up of
peptidoglycan); it is absent in animal cells.
• In plant cells, cell wall is composed of middle lamella, primary wall and secondary wall.
• Middle lamella is made up of calcium and magnesium pectate and acts as a cementing
layer between two adjoining cells.
• Primary wall lies inner to middle lamella and is composed of parallel fibres of cellulose
(polysaccharide) running through a matrix of other polysaccharides such as
hemicellulose and pectin.
• Middle lamella and primary wall are present in all the plant cells whereas secondary
wall is present only in some specialized cells like collenchyma, xylem vessels,
sclerenchyma etc.
• Secondary wall lies inner to primary wall and is composed of cellulose (in
collenchyma), lignin (in xylem vessels and sclerenchyma), suberin (in endodermis of
roots) etc.
Cell wall
• Fine strands of cytoplasm called plasmodesmata are present within cell
walls that connect the cytoplasm of adjacent cells.
• Thus plasmodesmata are the cytoplasmic connections between cells,
running transversally through the walls.
• They allow the passage of substances between the adjoining cells.
Functions of cell wall
• Cell wall provides mechanical strength and support for individual cells and the plant
as a whole.
• Cell wall prevents cells from bursting under the pressure created by the osmotic
entry of water. Cellulose has a very high tensile strength (i.e. it is resistant to
stretching) and makes it possible for a cell to withstand large pressures develop
within it due to osmosis. It prevents undue expansion of the cell when water enters
by osmosis to compensate for the lack of contractile vacuole. This prevents bursting
of cells.
• Cell wall allows water to pass along it and contributes to the movement of water
through the plant by apoplast and symplast pathways.
• Epidermal cells often have a waterproof layer of waxy cutin on their outer walls that
helps to reduce water loss by evaporation.
(Note: We will study cellulose in detail in the chapter “Biological molecules” and
osmosis in detail in the chapter “Cell membrane and transport”.)
Cell surface membrane
• Cell membrane or bio-membrane is thin, elastic and partially permeable
membrane (allows selected substances to pass through it to limited extent).
Cell membrane is of two types.
⮚Cell surface membrane (plasma membrane): It forms the outermost
boundary of cytoplasm in both prokaryotic and eukaryotic cells. It also
controls the exchange of materials such as nutrients and waste products
between the cell and its environment
⮚Internal membranes: They form the outer boundary of cell organelles.
Nuclear membrane, mitochondrial membrane, chloroplast membrane, ER
membrane etc. are some internal membranes.
• Cell membrane is basically composed of phospholipids and proteins.
Functions of cell surface membrane
• It separates the cell from its external environment and also controls
movement of substances in and out of cells.
• It is important in cell recognition.
• It has receptor sites which bind specific hormones and
neurotransmitters.
• In animal cells, it may be folded to form microvilli to provide a larger
surface area for the absorption of substances.
• It has components that attach to one another and so help cells to form
tissues.