Reproduction
Reproduction
Reproduction is the ability of living organisms to produce new individual of their type or new
offspring. Reproduction is necessary for increase in number and perpetuation of life.
Reproduction is divided into two types:
1. Sexual reproduction
2. Asexual reproduction
Asexual Reproduction
This is a type of reproduction which only one part is involved to produce offspring by itself without
formation and fusion of gametes. Offspring which are identical (clones) to the parent cells are
always produced.
Xteristics of asexual reproduction
1. It involves a single parent.
2. There is no gamete formation or fertilization.
3. The whole process takes place in a small period of time.
4. There is limited variation (genetically similar offsprings)
REPRODUCTION IN FUNGI
Fungi reproduce sexually and/or asexually.
Perfect fungi reproduce both sexually and asexually, while imperfect fungi reproduce only
asexually.
Deuteromycetes (fungi imperfecti) reproduce asexually only e.g Alternaria, Colletotrichum,
Trichoderma. They produce conidia as their asexual spores.
Asexual spores formed in Fungi
1. Conidium (formed in conidiophore) e.g Ascomycetes
2. Sporangiospores (formed in Sporangium) e.g Phycomycetes (Zygomycetes and oomycetes)
Sporangiospores are either motile and naked- zoospores, or nonmotile and walled -
aplanospores.
3. Basidiospore (formed in Basidium) e.g Basidiomycetes
Sexual spores formed in Fungi
1. Ascospore (formed in ascus) e.g Ascomycetes (Penicillium, Aspergillus).
2. Basidiospore: produced in Basidium by basidiomycetes (Mushroom, Agaricus, Puccinia,
Ustilago).
3. Zygospore: formed in Sporangium of Zygomycetes (Rhizopus, Mucor)
4. Oospore: formed in Oogonium in Oomycetes (Phytophthora)
PLANT REPRODUCTION
Flower is the reproductive structure of flowering plants. It has the male and female sex organs and
is the most advanced reproductive structure in plant kingdom. Fertilization occurs in flowers and
produces seeds which germinate into new plants.
Structure of an ovule
A mature ovule consists of:
• An embryo sac: the important structure in the ovule that produces haploid female gametes. It
consists of:
An egg cell and two synergid cells
Two polar nuclei in the centre that fuse to form diploid nucleus
Three antipodal cells
• The nucellus: the tissue enclosed by integument and is nutritive in function.
• The integuments: protective sheaths that surround the nucellus covering the ovule
• The micropyle: a tiny opening into the ovule.
• Hilum: point of attachment of the ovule to the placenta.
Structure of an ovule
A mature ovule consists of:
• An embryo sac: the important structure in the ovule that produces haploid female gametes. It
consists of:
An egg cell and two synergid cells
The egg cell forms the zygote after fertilization and gives rise to a diploid embryo.
The synergid cells nourish the egg cell and give direction to the pollen tube after pollination
has occurred. They majorly function to give signals and guidance needed by the pollen tube
inside the embryo sac of the ovule. If synergid cells are absent, no fertilization would occur.
Two polar nuclei in the centre that fuse to form diploid nucleus called the secondary nucleus
or definitive nucleus.
This secondary nucleus (diploid) forms the nutritive tissue called endosperm in the double
fertilization process.
Three antipodal cells
Antipodal cells are situated at the opposite end of the micropyle and they supply nutrition to
the whole female gametophyte (major function is nutrition of the embryo).
• The nucellus: the tissue enclosed by integument and is nutritive in function.
• The integuments: protective sheaths that surround the nucellus covering the ovule
• The micropyle: a tiny opening into the ovule.
• Hilum: point of attachment of the ovule to the placenta.
Classification of flowers
POLLINATION
Pollination is the transfer of mature pollen grains from the anthers of one flower to the mature
stigma of the same flower or another flower of the same plant or another plant of the same species.
Pollination precedes fertilization.
Types of pollination
1. Self pollination
Cross pollination: transfer of mature pollen grains from the anther of a flower to the stigma of a
flower of another plant of the same or closely related species e.g morning glory, hibiscus, pride of
Barbados. Cross pollination occurs between different plants. Cross pollination leads to the
production of healthier offspring than self pollination as offspring are more adapted to the
environmental conditions, it produces viable seeds and new varieties of species with desirable
characteristics. But the shortcomings of this type of pollination is that it relies on external agents
such as wind and insects whose presence at the right time cannot be guaranteed. It also causes
wastage of pollen grains.
Conditions that aid cross pollination
• Unisexuality in dioecious plants; higher carpels than stamens in monoecious plants.
• Dichogamy: ripening of anther and filament of a bisexual flower or flowers in the same plant
at different times.
Protandry: ripening of anthers before stigmas. Sunflower, okra, cotton
Protogyny: ripening of stigmas before anthers. Palms, figs
• Self incompatibility: if pollen grains from the same flower or flowers from the same plant
fall on the stigmas of some flowers, they wither and die. Thus, only pollen grains from another
plant can fertilize such plants. e.g passion flower and tea.
Pollinating agents
Wind, insect (major pollinators); water, other animals.
Flowers pollinated by insects are called entomophilous flowers and have the following
characteristics:
• They have large and conspicuous petals/ sepals with bright colours.
• They usually possess scents and nectar is also present.
• Pollen grains are rough, sticky and relatively few.
• Stigma is flat with sticky surface to receive pollens
• Petals are shaped and arranged to enable visiting insects become dusted with pollen grains.
• Short and pendulous filament
Examples of entomophilous flower: Hibiscus, Delonix, cowpea, crotolaria, pride of Barbados e.t.c.
Structure of an ovule
A mature ovule consists of:
• An embryo sac: the important structure in the ovule that produces haploid female gametes. It
consists of:
An egg cell and two synergid cells
The egg cell forms the zygote after fertilization and gives rise to a diploid embryo.
The synergid cells nourishes the egg cell and give direction to the pollen tube after pollination
has occurred. They majorly function to give signals and guidance needed by the pollen tube
inside the embryo sac of the ovule. If synergid cells are absent, no fertilization would occur.
Two polar nuclei in the centre that fuse to form diploid nucleus called the secondary nucleus
or definitive nucleus.
This secondary nucleus (diploid) forms the nutritive tissue called endosperm in the double
fertilization process.
Three antipodal cells
Antipodal cells are situated at the opposite end of the micropyle and they supply nutrition to
the whole female gametophyte (major function is nutrition of the embryo).
• The nucellus: the tissue enclosed by integument and is nutritive in function.
• The integuments: protective sheaths that surround the nucellus covering the ovule
• The micropyle: a tiny opening into the ovule.
• Hilum: point of attachment of the ovule to the placenta.
Endosperm
The primary endosperm nucleus divides many times to produce endosperm tissue.
It functions for food storage for the embryo.
It may remain as endosperm providing food for some embryos such as maize and castor plants. Or
transfer the function of food reserve to cotyledons and disappear as the cotyledons develop as in
the embryos of cowpea seeds.
Monocotyledonous seed: one seed leaf or cotyledon e.g rice, wheat, maize, sorghum.
Dicotyledonous seed: two seed leaves or cotyledons e.g beans, mango.
Not all seeds have endosperm. Seeds with endosperm are called endospermic seeds. Only
Angiosperms have endosperm (food reserve). E.g maize, wheat, castor bean, coconut, barley,
rubber.
GERMINATION OF SEEDS
Germination is the gradual development of the embryo of the seed into a seedling or a young plant.
Seeds pass through a period of rest called dormancy. Under unfavourable conditions, seeds can
withstand and remain alive (dormancy) but under favourable conditions, they can still germinate
(viable seeds).
Types of germination
• Epigeal germination: cotyledons are carried above the soil surface. seen in dicots e.g
cowpea, melon, mango, groundnut.
• Hypogeal germination: cotyledons remain in the soil surface. Seen in monocots e.g millet,
wheat, oil palm, maize.
FRUITS
A fruit is a mature fertilised ovary of a flower and contains one or more seeds.
Fruits that develop without fertilization are called Parthenocarpic fruits and are often seedless e.g
bananas and pineapples.
Structure of a fruit
1. Pericarp: fruit wall (develops from ovary wall). There are three layers of the pericarp:
a. Epicarp: outer covering
b. Mesocarp: middle layer
c. Endocarp: inner layer. Seeds are within the endocarp.
The pericarp may be soft, hard, dry or fibrous.
2. Seeds: seeds are arranged in the fruit just as ovules are arranged in the ovary. A seed has
hilum (a scar left by the point of attachment of the funicle of the seed to the placenta). An
embryo is present in each seed. Some seeds store food in endosperm while other store food in
cotyledons.
3. Fruit stalk: a scar showing point of attachment of the fruit to the plant due to receptacle of
the flower.
4. Remains of the style: a scar showing point of attachment of the fruit (developed from ovary)
to the style.
Differences between seed and fruit
• A fruit develops from ovary, has pericarp and contains seeds while a seed develops from
ovule, has seed coat and contains an embryo.
• A seed has micropyle (an opening), a fruit does not have a micropyle.
• A fruit has remains of style while a seed does not.
• A fruit has fruit stalk due to receptacle while a seed has a scar (hilum) formed by attachment
of funicle.
Types of fruits
A. Whether it develops from only ovary or not:
• True fruit: develops solely and from fertilized ovary e.g mango, orange and cowpea.
• False fruit: develops from an ovary and other floral parts e.g apple, pineapple, cashew.
B. Number of ovaries and flowers that form the fruit
• Simple fruit: develops from a single flower with a single ovary. Pistil maybe monocarpous
(cowpea, maize) or synocarpous (okra, tomato, pawpaw).
• Aggregate fruit: develops from a single flower with several ovaries; apocarpous pistil (pistil
with separate carpels). Each ovary develop into a fruitlet like in grapes and the fruitlets are
embedded in a fleshy receptacle.
* Flowers are solitary in simple and aggregate fruits.
• Composite fruit: develops from an inflorescence (group of flowers) or flowers that are close
to one another. All the fruitlets from each flower and the floral parts (as in false fruits)
develop into one big fruit. e.g pineapple and breadfruit.
C. Nature of pericarp (dry or fleshy)
• Fleshy fruits: with fleshy (thick, soft, succulent) pericarp. The fleshy parts of some fruits may
develop from other floral parts and may not include the pericarp.
Types of fleshy fruits: drupe, berry, pome, sorosis, hesperidium, syconium.
• Dry fruits: with dry, hard, woody or fibrous pericarp.
Types of dry fruits: dry dehiscent and dry indehiscent fruits.
**Dehiscence: when pericarp split when ripe to release seeds.
**All fleshy fruits are indehiscent.
Fleshy fruits
1. Drupe: true simple fruit with well developed pericarp. The pericarp of drupe has three layers:
Dry fruits
1. Dry dehiscent fruits: dry fruits that split to release the seeds when ripe.
Types of dry dehiscent fruits
a. Legume: simple fruit from a superior monocarpous pistil; has marginal placentation; fruit
dehisces longitudinally along both sides. e.g cowpea, soya beans, pride of Barbados,
flamboyant, crotolaria.
b. Follicle: simple fruit from a superior monocarpous pistil; ripe fruit dehisces longitudinally on
one side only e.g silk cotton, kola.
c. Capsule: simple fruit from a single ovary divided into many carpels; dehisces along many
sides. e.g castor oil fruit, para-rubber fruit, okra fruit and cotton.
d. Schizocarp: formed from an ovary made up of one carpel; breaks up into units each enclosing
one seed. e.g desmodium, cassia.
1. Dry indehiscent fruits: dry fruits that do not split when ripe; they fall to the ground, pericarp
decays to release seeds instead.
Types of dry indehiscent fruits
a. Caryopsis: simple fruit from monocarpous pistil; pericarp and seed coat are fused to form a
covering over seed. e.g maize grain, rice, millet, guinea corn.
b. Nut: true simple with a hard woody pericarp and has only one seed. e.g cashew nut
c. Achene: true fruit with only one seed from an apocarpous pistil. Pericarp is free from seed
coat e.g strawberry, sunflower.
d. Cypsela: simple true fruit from a superior monocarpous ovary and has only one seed;
pericarp is free from seed coat. e.g tridax fruit, emilia.
e. Samara: simple true fruit from superior ovary with > 1 carpels; the pericarp form wing like
structures. e.g combretum, pterocarpus (African rose flower).
Female Gametes
The female sex cells or gametes called the eggs or ova are produced in the ovaries by a process
called oogenesis. The human female gametes are larger than the sperms. Each ovum is about 0.1mm
in diameter. It consists of the cytoplasm, a nucleus in the centre, granules and yolk droplets. The
yolk provides a source of nourishment for the embryo, especially at the early stages of
development.
The ovum's cytoplasm is surrounded by two membranes. The outer one is called the vitelline
membrane while the inner one is the plasma membrane. The ovum is surrounded on the outside by
a jelly coat of variable thickness made of glycoprotein.
The nuclei of the sperm and ovum contain chromosomes (thread-like materials) which carry the
genes. The genes are responsible for the transmission of characteristics from parents to offspring.
When a female is born, her ovaries contain all potential egg cells or primary oocytes which are
about 20000 in each ovary. Only about 400-500 of these mature in a female's active reproductive
life.
Menstrual cycle
Maturation of the ovum occurs in one of the ovaries.
Several primary oocytes each start to develop in a structure called follicle. The cells surrounding
the oocyte in the Follicle continue to divide as the follicle matures.
Developing follicles secrete oestrogen.
Usually, only one of the developing follicles become mature and becomes a Graafian follicle and
contains a secondary oocyte. The matured ovum is then released into the Fallopian tube (a process
called ovulation).
There is thickening of the uterine endometrium after the release and the thickening is maintained
in preparation to receive a fertilized egg.
The remaining part of the Graafian follicle develops into corpus luteum (a solid yellow body which
secretes progesterone).
If the ovum does not become fertilized in the oviduct, it goes into the uterus where the uterine
endometrial tissue breaks down and are released to the exterior through the vaginal opening.
This is called menstruation.
The menstrual cycle is a 4-week cycle (about 28 days).
Functions of hormones in menstrual cycle
• FSH: starts menstrual cycle by stimulating follicle development.
• Oestrogen: causes the endometrium to build up and thicken. High amount of oestrogen in the
blood stimulate the release of LH.
• LH: stimulates ovuluation and development of corpus luteum from the Graafian follicle.
• Progesterone (pregnancy hormone): inhibits FSH and maturation of oocytes. Prevents
uterine contractions during pregnancy and aids development of placenta.
Oestrogen and progesterone maintain the endometrium. If the ovum does not fertilize, corpus
luteum shrivels and since there's no more progesterone to maintain the endometrial tissue, it
breaks down. In the absence of progesterone, FSH starts work again and the whole cycle is
repeated.
Male Gametes
The male sex cells or gametes called sperms are produced in the testes by a process called
spermatogenesis. The sperm or spermatozoon consists of a head which contains the nucleus, a
middle piece and a whip-like tail or flagellum. A human sperm is about 0.05mm long and
microscopic and is usually smaller than the egg (ovum).
Spermatogenesis
About 300 million sperms are produced daily by the seminiferous tubules in the testes.
The actively dividing germ cells in the seminiferous tubules undergo meiosis to produce immature
spermatids which develop into the spermatozoa. The sperm cells (spermatozoa) are carried to the
epididymis where they're stored and become mature.
The best temperature for sperm production is a little lower than body temperature (35°C). This is
why the testes descend into the scrotal sac (scrotum) outside the body.
The Leydig (interstitial) cells in the seminiferous tubules of the testes produce testosterone
responsible for the development of male sexual characters.
Hormonal function in male sexual activity
• FSH stimulate sperm production.
• LH which is called ICSH (interstitial cells stimulating hormone) in males aids testosterone
production by stimulating the interstitial cells (Leydig cells).
Fertilization in man
When being sexually aroused, blood flows into the genitals and the penis become erect in males
while the vagina produces a lubricating mucus to aid free thrusting of the penis into the vagina.
Ejaculation (release of semen) into the vagina gives makes a pleasant sensation called orgasm. The
female may also experience orgasm.
Each ejaculation release about 200-300 million sperms which can stay up to 3 -5 days in females
before dying. Fertilization can only occur if an egg cell still remains in the upper part of the oviduct
(about 24 hours after release from the ovary).
This means that fertilization can occur if sexual intercourse has taken place about 5 days before
ovulation and about a day after ovulation.
This means that usually, there are six possible days for fertilization in a human female per month.
When a sperm meets an egg cell, the acrosomal lytic enzymes break down the cell coats and
membranes of the egg cell and penetrate into it with its head leaving it's tail behind.
The nucleus of the sperm fuses with the egg nucleus to form diploid zygote. When a sperm enters
the egg cell, the zona pellucida prevents other sperms from entering. Out of the millions of sperms
ejaculated, only 1 fertilizes an ovum. When the ovum becomes fertilized, the corpus luteum
continue producing progesterone and oestrogen.
Fertilization takes place in the upper part of the oviduct.
If more than one egg is released and fertilized, the pregnancy involves more than one fetus, usually
two (twins). Because the genetic material in each egg and in each sperm is slightly different, each
fertilized egg is different. The resulting twins are thus fraternal twins. Identical twins result when
one fertilized egg separates into two embryos after it has begun to divide. Because one egg was
fertilized by one sperm, the genetic material in the two embryos is the same.
Development in humans
The zygote undergoes mitotic divisions and increases in size to form the embryo. The embryo
becomes implanted in the uterine wall. Uterus grows to compliment the growth of the embryo. The
embryonic membranes (amnion and chorion) develop. The amnion forms a sac containing amniotic
fluid. The embryo eventually develops into a foetus.
The placenta is a tissue containing both maternal and embryonic tissues. It is connected to the
foetus via the umbilical cord.
The foetal capillaries lie in finger-like projections called villi in the placenta and stick into the
maternal blood spaces. Here there is exchange materials between the mother and the foetus
without mixing of their blood. Trophoblast (a thin barrier between them) allows selective exchange
of materials). Oxygen and nutrients diffuse into the foetal blood from the maternal blood while
carbondioxide and urea from the foetal blood enter the maternal blood.
Gestation period in man is about 38 weeks.
• Uterus: implantation and development of the embryo occurs here
• Placenta: connection between mother and foetus for exchange of materials.
METAMORPHOSIS IN INSECTS
Metamorphosis is the gradual changes that occur during the development of an animal from the
zygote (fertilized egg) to the adult stage. There are two types of metamorphosis in insects which
include:
1. Complete metamorphosis (Holometabolism)
2. Incomplete metamorphosis (Hemimetabolism)
Complete metamorphosis: this is the series of gradual changes which takes place in insects from
fertilized eggs to larva then to pupa and finally to the adult stage:
Egg ——– larva ——- pupa ——- adult
No resemblance between pupa and adult stage. Insect undergoing complete metamorphosis include
housefly, mosquito, butterfly, moth, bees, wasps and beetles.
Incomplete metamorphosis: In this type of metamorphosis, the egg hatches into a nymph which
generally resembles the adult except that it is smaller, wingless and sexually immature e.g
grasshopper, cockroach, dragonfly, locust, aphids, termites, praying mantis.