Gorytes

Gorytes laticinctus, copyright Jeremy Early.

Belongs within: Gorytini.

Gorytes is a genus of medium-sized to large wasps found in Eurasia, North America and Africa that provision their nests with leafhoppers (Auchenorrhyncha). Nests (where known) are burrows about 12 cm deep with up to four cells per burrow (Bohart & Menke 1976).

Characters (from Bohart & Menke 1976): Medium-sized to fairly large wasps; inner eye margins converging below, often strongly; mandible usually with two inner subteeth; labrum inconspicuous; male flagellomeres often with tyli but last four articles not specially modified; pronotal collar rather thin, rounded, nearly even all across, not closely adherent to scutum; female foretarsal rake well developed, fore basitarsus with two long setae before apex; female arolia usually nearly equal in size; posterolateral oblique scutal carina present; prescutellar sulcus foveate; episternal sulcus when well impressed ending on scrobal sulcus which is continued forward in nearly a straight line to omaulus; acetabular carina incomplete or obscure; sternaulus complete or essentially so; forewing costal margin gently or sometimes strongly bowed out near base, media diverging before cu-a, stigma moderate, veinlet of submarginal cell II between recurrents about one-fourth posterior length of submarginal cell I; jugal lobe considerably larger than tegula, hindwing media generally diverging before cu-a but in some species at or a little beyond it; midtibia with two apical spurs; propodeal enclosure either extensively sculptured or longitudinally ridged, ridges sometimes confined to a frontal strip; spiracular groove present, usually distinct; gaster tapering toward base, sometimes nearly pedunculate; male with seven terga normally exposed, sterna without fimbriae, sternum VIII narrowly blade shaped, often minutely notched at apex; female pygidial plate variable, flat, usually long.

Gorytes Latreille 1804 (see below for synonymy) BM76
    |--*G. quinquecinctus (Fabricius 1793) BM76 (see below for synonymy)
    |--G. africanus Mercet 1905 BM76
    |--G. aino Tsuneki 1963 BM76
    |--G. albidulus (Lepeletier 1832) (see below for synonymy) BM76
    |--G. albosignatus Fox 1892 BM76
    |--G. ambiguus Handlirsch 1888 BM76
    |--G. angustus (Provancher 1895) [=Hoplisus angustus; incl. H. angustatus Ashmead 1899] BM76
    |--‘Hoplisus’ archoryctes Cockerell 1922 BM76
    |--G. atricornis Packard 1867 (see below for synonymy) BM76
    |--G. atrifrons Fox 1892 BM76
    |--G. browni GS61
    |--G. canaliculatus Packard 1867 (see below for synonymy) BM76
    |--G. cochisensis Bohart 1971 BM76
    |--G. cribratus Morawitz 1892 BM76
    |--G. deceptor Krombein 1958 BM76
    |--G. dorothyae Krombein 1950 BM76
    |    |--G. d. dorothyae BM76
    |    `--G. d. russeolus Krombein 1954 BM76
    |--G. eous Gussakovskij 1933 BM76
    |--G. fallax Handlirsch 1888 BM76
    |--G. flagellatus Bohart 1971 BM76
    |--G. flaviventris Morawitz 1894 BM76
    |--G. foveolatus Handlirsch 1888 (see below for synonymy) BM76
    |--G. hakutozanus Tsuneki 1963 BM76
    |--G. harbinensis Tsuneki 1967 BM76
    |--G. hebraeus Beaumont 1953 BM76
    |--G. heptapotamiensis Kazenas 1972 BM76
    |--G. intrudens Nurse 1903 BM76
    |--G. jonesi (Turner 1920) [=Arpactus jonesi] BM76
    |--G. koreanus Handlirsch 1888 BM76
    |--G. kulingensis (Yasumatsu 1943) [=Hoplisus kulingensis] BM76
    |--G. laticinctus (Lepeletier 1832) [=*Euspongus laticinctus] BM76
    |--G. limbellus Bohart 1971 BM76
    |--G. maculicornis (Morawitz 1889) [=Hoplisus maculicornis] BM76
    |--G. mcateei Krombein & Bohart 1962 BM76
    |--G. melpomene (Arnold 1936) [=Arpactus melpomene] BM76
    |--G. mongolicus Tsuneki 1970 BM76
    |--G. mystaceus MS01
    |--G. neglectus Handlirsch 1895 BM76
    |--G. nevadensis Fox 1892 BM76
    |--G. nigrifacies (Mocsáry 1879) [=Hoplisus nigrifacies] BM76
    |--G. pieli Yasumatsu 1943 BM76
    |--G. planifrons (Wesmael 1852) [=Hoplisus planifrons] BM76
    |--G. pleuripunctatus (Costa 1859) (see below for synonymy) BM76
    |    |--G. p. pleuripunctatus BM76
    |    `--G. p. barbarus Beaumont 1953 BM76
    |--G. procrustes Handlirsch 1888 BM76
    |--G. prosopis Bohart 1971 BM76
    |--G. provancheri Handlirsch 1895 (see below for synonymy) BM76
    |--G. proximus Handlirsch 1893 BM76
    |--G. quadrifasciatus (Fabricius 1804) (see below for synonymy) BM76
    |--G. quinquefasciatus (Panzer 1798) (see below for synonymy) BM76
    |    |--G. q. quinquefasciatus BM76
    |    `--G. q. levantinus Pulawski 1961 BM76
    |--G. radoszkowskyi Handlirsch 1888 BM76
    |--G. schlettereri Handlirsch 1893 BM76
    |    |--G. s. schlettereri BM76
    |    `--G. s. ponticus Beaumont 1967 BM76
    |--G. schmiedeknechtii Handlirsch 1888 BM76
    |--G. sepultus (Cockerell 1906) [=Hoplisus sepultus] BM76
    |--G. simillimus Smith 1856 [incl. G. ephippiatus Packard 1867, G. gyponacinus Rohwer 1911] BM76
    |--G. sogdianus Gussakovskij 1952 BM76
    |--G. sulcifrons (Costa 1869) [=Hoplisus sulcifrons; incl. H. laevigatus Kohl 1880] BM76
    |--G. takeuchii Tsuneki 1963 BM76
    |--G. tricinctus (Pérez 1905) [=Hoplisus tricinctus] BM76
    |--G. umatillae Bohart 1971 [incl. G. vicinus Handlirsch 1893 non Euspongus vicinus Lepeletier 1832] BM76
    `--G. verhoeffi Tsuneki 1967 BM76

Gorytes Latreille 1804 [=Euzonia Stephens 1829, Hoplisus Lepeletier 1832; incl. Arpactus Panzer 1805, Euspongus Lepeletier 1832] BM76

Gorytes albidulus (Lepeletier 1832) [=Hoplisus albidulus; incl. Euspongus albilabris Lepeletier 1832, Mellinus dissectus Panzer 1801 non Panzer 1801, Gorytes elegans Smith 1856 non Lepeletier 1832] BM76

Gorytes atricornis Packard 1867 [incl. G. decorus Fox 1895, Hoplisus elegantulus Smith 1908, G. rugosus Packard 1867] BM76

Gorytes canaliculatus Packard 1867 [incl. G. asperatus Fox 1895, Hoplisus corrugis Mickel 1918, G. geminus Handlirsch 1888] BM76

Gorytes foveolatus Handlirsch 1888 [incl. G. dichrous Mercet 1906, G. longicornis Handlirsch 1898 non Sphex longicornis Rossi 1790, G. rubrocinctulus Strand 1910, G. usurpator Schulz 1906] BM76

Gorytes pleuripunctatus (Costa 1859) [=Hoplisus pleuripunctatus; incl. G. fraternus Mercet 1906, H. tirolensis Kohl 1880] BM76

Gorytes provancheri Handlirsch 1895 [incl. G. laticinctus Provancher 1888 non Euspongus laticinctus Lepeletier 1832] BM76

Gorytes quadrifasciatus (Fabricius 1804) [=Mellinus quadrifasciatus, *Arpactus quadrifasciatus; incl. Hoplisus montivagus Mocsáry 1878, Euspongus vicinus Lepeletier 1832] BM76

*Gorytes quinquecinctus (Fabricius 1793) BM76 [=Mellinus quinquecinctus BM76, Ceropales quinquecinctus L02, *Euzonia quinquecincta BM76, *Hoplisus quinquecinctus BM76; incl. Mellinus arenarius Panzer 1798 BM76, Crabro calceatus Rossi 1794 BM76, Gorytes cinctus Latreille 1805 BM76, G. ruficornis Latreille 1805 BM76, Hoplisus sinuatus Costa 1869 BM76]

Gorytes quinquefasciatus (Panzer 1798) [=Mellinus quinquefasciatus; incl. Hoplisus anceps Mocsáry 1879, H. eburneus Chevrier 1870, H. geminatus Costa 1869, Gorytes intercedens Handlirsch 1893, H. lacordairei Lepeletier 1845, G. mauretanicus Handlirsch 1898] BM76

*Type species of generic name indicated

REFERENCES

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[GS61] Giordani Soika, A. 1961. Gli Odynerus sensu antiquo del continente Australiano e della Tasmania. Boll. Mus. Civ. Venezia 14: 57–202.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[MS01] Mocsáry, A., & V. Szépligeti. 1901. Hymenopterák [Hymenopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 121–169. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

Last updated: 30 December 2019.

Stizus

Stizus fasciatus, copyright Cor Zonneveld.

Belongs within: Stizini.

Stizus is a genus of medium-sized to large wasps found in Eurasia, North America and Africa that provision their nests with Orthoptera or mantids (Bohart & Menke 1976).

Characters (from Bohart & Menke 1976): Eyes converging slightly below, at least narrower below than above; a low and somewhat double convexity below mid-ocellus; mandible dentate subapically; exposed part of labrum moderately convex, usually a fourth to a third as long as broad; male antenna without a projection on flagellomere IX; scutellum typically (but not always) with a medial, pubescent depression in the female; propodeum not compressed into a ridge posterolaterally; midtibia with two apical spurs; forewing with submarginal cell II open in front; hindwing media diverging far before cu-a; median cell with two distal appendices; precoxal sulcus usually present.

<==Stizus Latreille 1802-1803 [incl. Megalostizus Schulz 1906, Megastizus Patton 1879, Stizolarra Saussure 1887] BM76
    |--S. ruficornis (Forster 1771) (see below for synonymy) BM76
    |    |--S. r. ruficornis BM76
    |    |--S. r. eremicus Bytinski-Salz 1955 BM76
    |    `--S. r. strigatus Mochi 1939 BM76
    |--S. adelphus Kirby 1900 BM76
    |--S. aegyptius Lepeletier 1845 BM76
    |--S. aestivalis Mercet 1906 BM76
    |    |--S. a. aestivalis BM76
    |    `--S. a. merceti Beaumont 1951 BM76
    |--S. anchoratus Mantero 1917 BM76
    |--S. anchorites Turner 1920 BM76
    |--S. annulatus (Klug 1845) (see below for synonymy) BM76
    |--S. apicalis Guérin-Méneville 1844 BM76
    |--S. arenarum Handlirsch 1892 BM76
    |--S. arnoldi Mochi 1939 BM76
    |--S. atrox (Smith 1856) [=Larra atrox; incl. L. flavomaculata Smith 1873, Stizus pentheres Handlirsch 1895] BM76
    |--S. aurifluus Turner 1916 BM76
    |--S. basalis Guérin-Méneville 1844 [incl. Larra infuscata Klug 1845] BM76
    |--S. baumanni Handlirsch 1901 BM76
    |--S. beccarii Mantero 1917 BM76
    |--S. bensoni Arnold 1951 BM76
    |--S. berlandi Arnold 1945 BM76
    |--S. biclipeatus (Christ 1791) (see below for synonymy) BM76
    |--S. bipunctatus (Smith 1856) [=Larra bipunctata] BM76
    |--S. bizonatus Spinola 1838 [incl. Larra sphegiformis Klug 1845] BM76
    |--S. brevipennis Walsh 1869 [=*Megastizus brevipennis; incl. Larra brendeli Taschenberg 1875] BM76
    |--S. breyeri Arnold 1936 BM76
    |--S. chrysorrhoeus Handlirsch 1892 BM76
    |--S. coloratus Nurse 1903 BM76
    |--S. combustus (Smith 1856) [=Larra combusta; incl. L. fuliginosa Klug 1845 non Dahlbom 1843] BM76
    |--S. continuus (Klug 1835) [=Larra continua] BM76
    |--S. delessertii Guérin-Méneville 1844 BM76
    |--S. deserticus Giner Marí 1945 BM76
    |--S. dewitzii Handlirsch 1892 [incl. S. erythraspis Cameron 1910, S. propodealis Bischoff 1915] BM76
    |--S. dispar Morawitz 1888 BM76
    |--S. elegans Dahlbom 1845 BM76
    |--S. ellenbergeri Arnold 1929 BM76
    |--S. emir Handlirsch 1901 BM76
    |--S. erythraeensis Mantero 1917 BM76
    |--S. erythrocephalus (Taschenberg 1880) [=Larra erythrocephala] BM76
    |--S. euchromus Handlirsch 1892 BM76
    |--S. eugeniae Gussakovskij 1935 BM76
    |--S. excellens Lohrmann 1943 [incl. S. eximius Arnold 1936 non Morawitz 1894] BM76
    |--S. eximius Morawitz 1894 BM76
    |--S. fasciatus (Fabricius 1781) (see below for synonymy) BM76
    |--S. fedtschenkoi Radoszkowski 1877 BM76
    |--S. ferrandii Magretti 1898 BM76
    |--S. ferrugineus (Smith 1856) [=Larra ferruginea; incl. S. ritsemae Handlirsch 1895] BM76
    |--S. franzi Turner 1916 [incl. S. congoensis Mantero 1917, S. neavei Kohl 1913 non Turner 1912] BM76
    |--S. fuscatus Bingham 1897 BM76
    |--S. fuscipennis (Smith 1856) [=Larra fuscipennis] BM76
    |--S. gobiensis Tsuneki 1970 BM76
    |--S. gracilipes Handlirsch 1892 BM76
    |--S. hamatus Arnold 1929 BM76
    |--S. handlirschi Radoszkowski 1893 BM76
    |--S. hebraeus Balthasar 1954 BM76
    |--S. hispanicus Mocsáry 1883 [incl. S. villosus Costa 1887] BM76
    |--S. histrio Morawitz 1888 BM76
    |--S. hugelii Handlirsch 1892 BM76
    |--S. hyalipennis Handlirsch 1892 BM76
    |--S. imperator Nurse 1903 BM76
    |--S. imperialis Handlirsch 1892 [incl. S. tulbaghensis Arnold 1936] BM76
    |    |--S. i. imperalis BM76
    |    `--S. i. conspicuus Arnold 1952 BM76
    |--S. iridis Dow 1942 BM76
    |--S. jordanicus Lohmann 1942 BM76
    |--S. kaszabi Tsuneki 1971 BM76
    |--S. kiseritzkii Gussakovskij 1928 BM76
    |--S. koenigi Morawitz 1888 [incl. S. tages Kirby 1889] BM76
    |--S. kohlii Mocsáry 1883 BM76
    |--S. lacteipennis Mocsáry 1883 BM76
    |--S. lepidus (Klug 1845) [=Larra lepida] BM76
    |--S. lineatus (Fabricius 1793) [=Bembex lineatus] BM76
    |--S. lohrmanni Bohart in Bohart & Menke 1976 (see below for synonymy) BM76
    |--S. lughensis Magretti 1898 BM76
    |--S. luteotaeniatus Gussakovskij 1933 BM76
    |--S. lutescens Radoszkowski 1877 BM786
    |--S. marnonis Handlirsch 1892 BM76
    |--S. marshalli Turner 1912 BM76
    |--S. marthae Handlirsch 1892 [incl. S. cheops Morice 1897, S. cheops Bingham 1897 non Morice 1897] BM76
    |--S. melanoxanthus (Smtih 1856) [=Larra melanoxantha] BM76
    |--S. melanurus Handlirsch 1892 BM76
    |--S. melleus (Smith 1856) [=Larra mellea] BM76
    |--S. mocsaryi Handlirsch 1895 BM76
    |--S. mongolicus Tsuneki 1971 BM76
    |--S. multicolor Turner 1916 BM76
    |--S. nadigi Roth 1933 BM76
    |--S. neavei Turner 1912 BM76
    |--S. nigriventris Arnold 1951 BM76
    |--S. niloticus Handlirsch 1892 BM76
    |--S. occidentalis Parker 1929 BM76
    |--S. ocellatus Gistel 1857 BM76
    |--S. orientalis Cameron 1890 BM76
    |--S. pauli Mantero 1917 BM76
    |--S. perrisii Dufour 1838 BM76
    |    |--S. p. perrisii BM76
    |    `--S. p. ibericus Beaumont 1962 BM76
    |--S. pictus Dahlbom 1845 BM76
    |--S. pluschtschevskii Radoszkowski 1888 BM76
    |--S. praestans Morawitz 1893 BM76
    |--S. pulcherrimus (Smith 1856) [=Larra pulcherrima] BM76
    |--S. pygidialis Handlirsch 1892 BM76
    |--S. quartinae (Gribodo 1884) [=Sphecius quartinae] BM76
    |--S. raddei Handlirsch 1889 BM76
    |--S. rapax Handlirsch 1892 BM76
    |--S. richardsi Arnold 1959 BM76
    |--S. rubellus Turner 1912 BM76
    |--S. rubroflavus Turner 1916 BM76
    |--S. rufescens (Smith 1856) [=Larra rufescens] BM76
    |--S. rufipes (Fabricius 1804) [=Liris rufipes; incl. S. gigas Walsh & Riley 1868, S. grandis Lepeletier 1845] BM76
    |--S. rufiventris Radoszkowski 1877 [incl. S. sarmaticus Morawitz 1891] BM76
    |    |--S. r. rufiventris BM76
    |    `--S. r. compar Handlirsch 1892 BM76
    |--S. rufocinctus Dahlbom 1845 BM76
    |--S. rufoniger Mochi 1939 BM76
    |--S. saharae Roth 1934 BM76
    |--S. saussurei Handlirsch 1895 BM76
    |--S. savignyi Spinola 1838 [incl. Larra succinea Klug 1845] BM76
    |--S. schmiedeknechti Handlirsch 1898 BM76
    |--S. scoliaeformis Arnold 1929 BM76
    |--S. scolioides Kokujev 1902 BM76
    |--S. sexfasciatus (Fabricius 1793) [=Bembex sexfasciatus] BM76
    |--S. spectrum Handlirsch 1901 BM76
    |--S. spinulosus Radoszkowski 1876 BM76
    |--S. stevensoni Arnold 1936 BM76
    |--S. storeyi Turner 1920 BM76
    |--S. tenuicornis (Smith 1856) [=Larra tenuicornis] BM76
    |--S. texanus Cresson 1872 BM76
    |--S. transcaspicus Radoszkowski 1893 BM76
    |--S. tricolor Handlirsch 1892 BM76
    |--S. tridens MS01
    |--S. tridentatus [=Bembex tridentata] L02
    |--S. tunetanus Costa 1893 BM76
    |--S. vespiformis (Fabricius 1775) [=Sphex vespiformis, *Stizolarra vespiformis] BM76
    |--S. vespoides (Walker 1871) [=Larra vespoides; incl. L. argentea Taschenberg 1875, L. magnifica Smith 1873] BM76
    |--S. wheeleri Arnold 1959 BM76
    |--S. zhelochovtzevi Gussakovskij 1952 BM76
    |--S. zimini Gussakovskij 1928 BM76
    |--S. zonatus (Klug 1845) [=Larra zonata; incl. L. dimidiata Taschenberg 1875 non Panzer 1806-1809] BM76
    `--S. zonosoma Handlirsch 1895 BM76

Stizus annulatus (Klug 1845) [=Larra annulata; incl. S. eversmanni Radoszkowski 1877, S. kisilkumii Radoszkowski 1877, S. picticornis Morawitz 1891, Larra subapicalis Walker 1871] BM76

Stizus biclipeatus (Christ 1791) [=Vespa biclipeata; incl. Larra cingulata Fabricius 1798, L. nubilipennis Smith 1856, Tiphia ruficornis Fabricius 1787 nec Vespa ruficornis Forster 1771 nec Bembex ruficornis Fabricius 1787] BM76

Stizus fasciatus (Fabricius 1781) [=Bembex fasciatus; incl. B. integra Fabricius 1804, Stizus terminalis Dahlbom 1845] BM76

Stizus lohrmanni Bohart in Bohart & Menke 1976 [=S. arnoldi Lohrmann 1943 non Mochi 1939; incl. S. pubescens Arnold 1929 non Klug 1835] BM76

Stizus ruficornis (Forster 1771) [=Vespa ruficornis; incl. S. distinguendus Handlirsch 1901, V. gaditana Gmelin 1790, S. komarovi Radoszkowski 1888, S. ornatus Dahlbom 1845, Larra pubescens Klug 1835, Bembex ruficornis Fabricius 1787 non Vespa ruficornis Forster 1771, *Stizus ruficornis, S. scutellaris Kirby 1900] BM76

*Type species of generic name indicated

REFERENCES

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[MS01] Mocsáry, A., & V. Szépligeti. 1901. Hymenopterák [Hymenopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 121–169. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.